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UNITED STATES
SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
____________________________
SCHEDULE 14A
Proxy Statement Pursuant to Section 14(a) of the
Securities Exchange Act of 1934
____________________________ 
Filed by the RegistrantFiled by a Party other than the Registrant 
Check the appropriate box:
 Preliminary Proxy Statement
 Confidential, for Use of the Commission Only (as permitted by Rule 14a-6(e)(2))
 Definitive Proxy Statement
 Definitive Additional Materials
 Soliciting Material Pursuant to §240.14a-12
Berkeley Lights,
PhenomeX Inc.
(Name of Registrant as Specified In Its Charter)

(Name of Person (s)Person(s) Filing Proxy Statement, if other than the Registrant)


Payment of Filing Fee (Check the appropriate box):
 No fee required.
 Fee computed on table below per Exchange Act Rules 14a-6(i)(1) and 0-11.
1)Title of each class of securities to which transaction applies:
2)Aggregate number of securities to which transaction applies:


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3)Per unit price or other underlying value of transaction computed pursuant to Exchange Act Rule 0-11 (set forth the amount on which the filing fee is calculated and state how it was determined):
4)Proposed maximum aggregate value of transaction:
5)Total fee paid:
Fee paid previously with preliminary materials.
Check box if any part of the fee is offset as providedFee computed on table in exhibit required by Item 25(b) per Exchange Act Rule 0-11(a)(2)Rules 14a-6(i)(1) and identify the filing for which the offsetting fee was paid previously. Identify the previous filing by registration statement number, or the Form or Schedule and the date of its filing.
1)Amount Previously Paid:
2)Form, Schedule or Registration Statement No.:
3)Filing Party:
4)Date Filed:0-11.




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March 31, 2021April 14, 2023
To Stockholders of Berkeley Lights,PhenomeX Inc.:
Our 20212023 Annual Meeting of Stockholders (“20212023 Annual Meeting”) will be held on Friday,Thursday, May 14, 202125, 2023 at 8:45 a.m.1:30 p.m. PDT. The 20212023 Annual Meeting will be conducted exclusively online via a live webcast. You will be able to attend the meeting, submit your questions during the meeting and vote your shares electronically at the meeting by visiting www.virtualshareholdermeeting.com/BLI2021.CELL2023. Because the meeting is completely virtual and being conducted via the Internet, stockholders will not be able to attend the meeting in person. The attached notice and proxy statement describe the formal business to be transacted at the meeting.
Whether or not you attend the virtual 20212023 Annual Meeting, it is important that your shares be represented and voted at the meeting. Therefore, I urge you to promptly vote and submit your proxy via regular mail, online or by telephone. The 20212023 Annual Meeting is being held so that stockholders may considerconsider:

Proposal 1: the election of Class IIII directors, and

Proposal 2: the ratification of the appointment of KPMG LLP as our independent registered public accounting firm for the fiscal year ending December 31, 2021.2023, and

Proposal 3: approval, on an advisory basis, of the compensation of our named executive officers.

Our board of directors determined that the matters to be considered at the 20212023 Annual Meeting are in the best interests of us and our stockholders. Forstockholders and unanimously recommends that you vote “FOR” the reasons set forthelection of each of the nominees named in Proposal 1 of the accompanying proxy statement, “FOR” the ratification of the appointment of KPMG LLP as our independent registered public accounting firm for the fiscal year ending December 31, 2023 in Proposal 2 and “FOR” the approval, on an advisory basis, of the compensation of our named executive officers in Proposal 3. Information about the matters to be acted upon at the 2023 Annual Meeting is contained in the Proxy Statement, the board of directors unanimously recommends a vote “FOR” each matter to be considered.accompanying proxy statement.

On behalf of the board of directors and the officers and employees of Berkeley Lights,PhenomeX Inc., I would like to take this opportunity to thank our stockholders for their continued support.
Sincerely,

erichobbssignatureforsecfia.jpgSincerely,
Eric Hobbs,siddharthasignaturea.jpg
Siddhartha Kadia, Ph.D.
Chief Executive Officer and Director


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Berkeley Lights,PhenomeX Inc.
NOTICE OF ANNUAL MEETING OF STOCKHOLDERS
Notice is hereby given that our 20212023 Annual Meeting of Stockholders (“20212023 Annual Meeting”) will be held virtually via live webcast on the Internet on Friday,Thursday, May 14, 2021, 202125, 2023 at 8:45 a.m.1:30 p.m. PDT for the following purposes:
1.Election of Class IIII directors;
2.Ratification of the appointment of KPMG LLP as our independent registered public accounting firm for the fiscal year ending December 31, 2021; and2023;
3.To transact suchApproval, on an advisory basis, of the compensation of our named executive officers; and
4.Such other businessmatters as may properly come before the meeting or any adjournment or postponement thereof.
These proposals are more fully described in the Proxy Statement following this Notice.Notice of Internet Availability of Proxy Materials (“Notice”).
Our board of directors recommends that you vote (i) FOR the election of the respective nominees for Class IIII directors named in the Proxy Statement to serve as directors of the Company andCompany; (ii) FOR the ratification of the appointment of KPMG LLP as our independent registered public accounting firm for the fiscal year ending December 31, 2021.2023; and (iii) FOR the approval, on an advisory basis, of the compensation of our named executive officers. You will be able to attend the meeting, submit your questions during the meeting and vote your shares electronically at the meeting by visiting www.virtualshareholdermeeting.com/BLI2021.CELL2023.
Our board of directors has fixed the close of business on March 23, 202130, 2023 as the record date for the determination of the stockholders entitled to notice of, and to vote at, the 20212023 Annual Meeting. Accordingly, only stockholders of record at the close of business on that date will be entitled to vote at the 20212023 Annual Meeting.
YOUR VOTE IS IMPORTANT. Whether or not you plan to attend the 20212023 Annual Meeting, we urge you to submit your vote via regular mail, online or via telephone as soon as possible to ensure your shares are represented. For additional instructions for each of these voting options, please refer to the proxy card. We encourage you to vote by internetInternet or telephone. It is convenient, conserves paper and saves the Company postage and other costs. Returning the proxy does not deprive you of your right to attend the 20212023 Annual Meeting and to vote your shares at the 20212023 Annual Meeting. TheThis Proxy Statement explains proxy voting and the matters to be voted on in more detail.
Important Notice Regarding the Availability of Proxy Materials for the 20212023 Annual Meeting to Be Held on May 14, 2021.25, 2023. Our proxy materials, including the Proxy Statement and Annual Report to Stockholders, are being made available on or about March 31, 2021April 14, 2023 on our website at https://investors.berkeleylights.cominvestors.phenomex.com in the Financial Information/SEC FilingsFinancials section of our Investors web page,Relations webpage, as well at the following website: www.proxyvote.com. We are providing access to our proxy materials over the Internet under the rules adopted by the U.S. Securities and Exchange Commission.

By order of the board of directors,
erichobbssignatureforsecfia.jpgsiddharthasignaturea.jpg
Eric Hobbs,Siddhartha Kadia, Ph.D.
Chief Executive Officer and Director
March 31, 2021April 14, 2023
Your vote is important. To vote your shares, please follow the instructions in the Notice of Internet Availability of Proxy Materials, which is being mailed to you on or about April 1, 2021.14, 2023.
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BERKELEY LIGHTS, INC.PhenomeX Inc.
PROXY STATEMENT
ANNUAL MEETING OF STOCKHOLDERS
May 14, 2021
JOBS ACT EXPLANATORY NOTE
We are an “emerging growth company” under applicable federal securities laws and are therefore permitted to take advantage of certain reduced public company reporting requirements. As an emerging growth company, we provide in this proxy statement the scaled disclosure permitted under the Jumpstart Our Business Startups Act of 2012 (the “JOBS Act”), including the compensation disclosures required of a “smaller reporting company,” as that term is defined in Rule 12b-2 promulgated under the Securities Exchange Act of 1934, as amended (the “Exchange Act”). In addition, as an emerging growth company, we are not required to conduct votes seeking approval, on an advisory basis, of the compensation of our named executive officers or the frequency with which such votes must be conducted. We will remain an “emerging growth company” until the earliest of (i) December 31, 2025, (ii) the last day of the fiscal year in which our annual gross revenues are $1.07 billion or more, (iii) the date on which we have, during the previous rolling three-year period, issued more than $1 billion in non-convertible debt securities or (iv) the date on which we are deemed to be a “large accelerated filer” as defined in the Exchange Act.25, 2023
IMPORTANT INFORMATION REGARDING DELIVERY OF PROXY MATERIALS
The Securities and Exchange Commission has adopted rules regarding how companies must provide proxy materials to their stockholders. These rules are often referred to as “notice and access,” under which a company may select either of the following options for making proxy materials available to its stockholders:
the full set delivery option; or
the notice only option.
A company may use a single method for all of its stockholders or use full set delivery for some while adopting the notice only option for others. In connection with our 20212023 Annual Meeting of Stockholders (the “2021(“2023 Annual Meeting”), we have elected to use the notice only option.
Full Set Delivery Option
Under the full set delivery option, which we have elected NOTnot to use for the 20212023 Annual Meeting, a company delivers all proxy materials to its stockholders by mail as it would have done prior to the change in the rules. In addition to delivery of proxy materials to stockholders, the company must post all proxy materials on a publicly-accessible website and provide information to stockholders about how to access the website. Although we have elected to use the notice only option in connection with the 20212023 Annual Meeting, we may choose to use the full set delivery option in the future.
Notice Only Option
Under the notice only option, a company must post all proxy materials on a publicly-accessible website. Instead of delivering proxy materials to its stockholders, the company instead delivers a “Notice of Internet Availability of Proxy Materials.”Materials” (“Notice”). The noticeNotice includes, among other things:

information regarding the date and time of the 2021 Annual Meetingannual meeting of stockholders as well as the items to be considered at the meeting;
information regarding the website where the proxy materials are posted; and
various means by which a stockholder can request paper or e-mail copies of the proxy materials.
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By reducing the amount of materials that a company needs to print and mail, the notice only option provides an opportunity for cost savings as well as conservation of paper products. As stated above, in connection with our 20212023 Annual Meeting, we have elected to use the notice only option. Accordingly, you will not receive all proxy materials by mail. These proxy materials include the Notice of 2021relating to the 2023 Annual Meeting, of Stockholders, proxy statement,Proxy Statement, proxy card and our Annual Report on Form 10-K, which are available at www.proxyvote.com. If a stockholder requests paper copies of the proxy materials, these materials must be sent to the stockholder within three business days and by first class mail.

ABOUT THE MEETING
Our 20212023 Annual Meeting will be held on Friday,Thursday, May 14, 2021,25, 2023, at 8:45 a.m.1:30 p.m. PDT. This year’s annual meeting will be a virtual meeting via live webcast on the Internet. You will be able to attend the 20212023 Annual Meeting, vote and submit your questions during the meeting by visiting www.virtualshareholdermeeting.com/BLI2021.CELL2023. In order to vote or submit a question during the 20212023 Annual Meeting, you will need to follow the instructions posted at www.proxyvote.com and will need the control number included onin your Notice or proxy card.Notice. If you do not have a control number,
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you will be able to listen to the meeting only. Youonly, and you will not be able to vote or submit your questions during the meeting.
Why am I receiving these materials?
Our board of directors is providing these proxy materials to you in connection with the solicitation by our board of directors of proxies for use at our 20212023 Annual Meeting, which will take place on May 14, 2021.25, 2023. Stockholders are invited to attend the 20212023 Annual Meeting and are requested to vote on the proposals described in this proxy statement (“Proxy Statement.Statement”).
All stockholders will have the ability to access the proxy materials via the Internet, including this Proxy Statement and our Annual Reportannual report on Form 10-K for the fiscal year ended December 31, 2020 (the “Annual2022 (“Annual Report”), as filed with the U.S. Securities and Exchange Commission (the “SEC”(“SEC”) on March 12, 2021.February 23, 2023. The Notice includes information on how to access the proxy materials, how to submit your vote over the Internet, by phone or by mail, as well as how to request a paper copy of the proxy materials. This Proxy Statement and the Annual Report are available at www.proxyvote.com.
What is the purpose of the 20212023 Annual Meeting?
At our 20212023 Annual Meeting, stockholders will act upon the matters outlined in the notice of meeting on the cover page of this proxy statement,Proxy Statement, consisting of 1) the election of Class IIII directors; 2) the ratification of the appointment of KPMG LLP as our independent registered public accounting firm for the fiscal year ending December 31, 2021;2023; 3) the approval, on an advisory basis, of the compensation of our named executive officers; and 3) any4) such other matters thatas may properly come before the meeting.meeting or any adjournment or postponement thereof.
Who is entitled to vote at the 20212023 Annual Meeting?
Only holders of our common stock as of the close of business on March 23, 2021,30, 2023, the record date, are entitled to receive notice of and to vote at the 20212023 Annual Meeting. In deciding all matters at the 20212023 Annual Meeting, each stockholder will be entitled to one vote for each share of our common stock owned as of the record date. We do not have cumulative voting rights for the election of directors. As of the record date, there were 66,278,21398,744,915 shares of our common stock outstanding and entitled to vote. We do not have any outstanding shares of preferred stock. A list of stockholders entitled to vote at the meeting will be available for examination during the meeting to any stockholder for any purpose germane to the meeting.
If, at the close of business on March 23, 2021,30, 2023, your shares of common stock were registered directly in your name with American Stock Transfer & Trust Company, LLC, our transfer agent, then you are a stockholder of record. As a stockholder of record, you may vote online at the 20212023 Annual Meeting or vote by proxy by completing, dating and signing the proxy card that was included with the proxy statement and promptly returning it in the preaddressed, postage paid envelope provided to you, or by submitting a proxy over the Internet or by telephone by following the instructions on the Notice. You may access instructions set forth in the Notice, proxy card.
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materials and our Annual Report to stockholders at www.proxyvote.com.
If your shares of common stock are held in street name through a broker, bank or other nominee, you are considered the beneficial owner of those shares and you have the right to instruct your broker, bank or other nominee, who is considered the stockholder of record for the purposes of voting at the 20212023 Annual Meeting, on how to vote the shares in your account. Your broker, bank or nominee will send you a voting instruction form for you to use to direct how your shares should be voted.
Brokers or other nominees who hold shares of common stock in “street name” for a beneficial owner of those shares typically have the authority to vote in their discretion on “routine” proposals when they have not received instructions from beneficial owners. However, brokers are not allowed to exercise their voting discretion with respect to the election of directors or for any matter that the SEC determines to be a “significant matter” without specific instructions from the beneficial owner. These non-voted shares are referred to as “broker non-votes.” If your broker holds your common stock in “street name,” your broker is not entitled to vote your shares on “non-routine” proposals (i.e., of the proposals to be considered at the 20212023 Annual Meeting, the election of directors)directors and “say-on-pay” votes) without your instructions and will only vote your shares on such proposals if you provide instructions on how to vote by filling out the voter instruction form sent to you by your broker with this proxy statement.Proxy Statement. Broker “non-votes”
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“non-votes” are not included in the tabulation of voting results for “non-routine” proposals for purposes of determining whether such proposals have been approved.
What constitutes a quorum?
The presence at the meeting or representation by proxy of the holders of common stock representing a majority of the voting power of the issued and outstanding shares of stock on the record date will constitute a quorum, permitting the meetingcompany to conduct its business.business at the meeting. Virtual attendance at our 2023 Annual Meeting constitutes presence in person for purposes of a quorum at the meeting. Each of (a) the shares held by each stockholder who properly submits a proxy, (b) the shares represented by broker non-votes that are present in person or represented by proxy and entitled to vote at the 2023 Annual Meeting and (c) the votes to ABSTAIN will be counted for purposes of determining the presence of a quorum at the 2023 Annual Meeting. Proxies received but marked as abstentions and broker non-votes will be included in the calculation of the number of shares considered to be present at the meeting for purposes of determining a quorum. If less than a majority of the combined voting power of the outstanding shares of common stock is represented at the 20212023 Annual Meeting, a majority of the shares so represented may adjourn the 20212023 Annual Meeting from time to time without further notice.
What vote is required to approve each item?
For the Election of Class IIII Directors. For purposes of electing directors at the 20212023 Annual Meeting, directors shall be elected by a plurality of the votes cast (meaning that the two director nominees who receive the highest number of shares voted “for” their election are elected). With respect to the election of directors, you may vote “for” or “withhold” authority to vote for each of the nominees for the Board. “Withhold” votes and broker non-votes are not considered votes cast for the foregoing purpose, and will have no effect on the election of the nominees.
For the Ratification of Independent Registered Public Accounting Firm. The affirmative vote of theholders of a majority in voting power of the votes cast (meaning the number of shares voted “for” the proposal must exceed the number of shares voted “against” such proposal) is required for the ratification of the appointment of KPMG LLP and approval of any other matter that may be submitted to a vote of our stockholders.ratification. You may vote “for,” “against” or “abstain” from voting on this proposal. Abstentions and broker non-votes are not considered votes cast for the foregoing purpose, and will have no effect on the vote for this proposal.
For the Advisory Vote on the Compensation of our Named Executive Officers. The affirmative vote of holders of a majority in voting power of the votes cast is required for the approval. You may vote “for,” “against” or “abstain” from voting on this proposal. Abstentions are not considered votes cast for the foregoing purpose, and will have no effect on the vote for this proposal.
For the Approval of any Other Matter that may be Submitted to a Vote of our Stockholders. The affirmative vote of holders of a majority in voting power of the votes cast may be submitted to a vote of our stockholders.
The inspector of election for the 20212023 Annual Meeting shall determine the number of shares of common stock represented at the meeting, the existence of a quorum and the validity and effect of proxies and shall count and tabulate ballots and votes and determine the results thereof.
What are the Board of Directors’ recommendations?
As more fully discussed under the section titled “Matters to Come Before the 20212023 Annual Meeting,” our board of directors recommends a votevote; FOR the election of the respective nominees for the Class IIII directors named in this proxy statement andProxy Statement; FOR the ratification of the appointment of KPMG LLP.LLP; and FOR the approval, on an advisory basis, of the compensation of our named executive officers.
Unless contrary instructions are indicated on the enclosed proxy, all shares represented by valid proxies received pursuant to this solicitation (and which have not been revoked in accordance with the procedures set forth below)
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will be voted (1) FOR the election of the respective nominees for director named in this proxy statement;Proxy Statement; (2) FOR the ratification of the appointment of KPMG LLP; (3) FOR the approval, on an advisory basis, of the compensation of our named executive officers; and (3)(4) in accordance with the recommendation of our board of directors, FOR or AGAINST all other matters as may properly come before the 20212023 Annual Meeting. In the event a stockholder specifies a different choice by means of the enclosed proxy, such shares will be voted in accordance with the specification made.
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How do I vote?
You may vote by following the instructions set forth in theyour Notice or on your proxy card or, if you are a street name holder (that is, if you hold your shares through a bank, broker or other holder of record), you must vote in accordance with the voting instruction form provided by your bank, broker or other holder of record. You may access the Notice, proxy materials and our annual reportAnnual Report to stockholders at www.proxyvote.com. If you are a street name holder, the availability of telephone or internetInternet voting will depend upon your bank’s, broker’s or other holder of record’s voting process.

Can I change my vote after I return my proxy card?
Yes. The giving of a proxy does not eliminate the right to vote at the 20212023 Annual Meeting should any stockholder giving the proxy so desire. Stockholders have an unconditional right to revoke their proxy at any time prior to the exercise of that proxy, by voting at the 20212023 Annual Meeting, by filing a written revocation or duly executed proxy bearing a later date with our Corporate Secretary at our headquarters. Note that if your shares of our common stock are held in street name through a broker, bank, or other nominee, you should contact the record holder of your shares regarding how to revoke your proxy or change your vote.
How do I attend the virtual 20212023 Annual Meeting?
The 20212023 Annual Meeting will be held entirely online due to the public health impact of the coronavirus outbreak (COVID-19) and to support the health and well-being of our partners, employees and stockholders.online. Stockholders as of March 23, 202130, 2023 will be able to attend and participate in the 2023 Annual Meeting online by accessing www.virtualshareholdermeeting.com/BLI2021.CELL2023. To join the 20212023 Annual Meeting, you will need to have your control number which is included on your Notice and your proxy card.
Even if you plan to attend the 20212023 Annual Meeting online, we recommend that you also vote by proxy as described herein so that your vote will be counted if you decide not to attend the 20212023 Annual Meeting.
Access to the Audio Webcast of the Annual Meeting. The live audio webcast of the 20212023 Annual Meeting will begin promptly at 8:45 a.m.1:30 p.m., PDT. Online access to the audio webcast will open approximately 15 minutes prior to the start of the 20212023 Annual Meeting to allow time for you to log in and test the computer audio system. We encourage our stockholders to access the meeting prior to the start time.
Log in Instructions. To attend the online 20212023 Annual Meeting, log in at www.virtualshareholdermeeting.com/BLI2021.CELL2023. Stockholders will need their unique control number which appears on the Notice (printed in the box and marked by the arrow) and the instructions that accompanied the proxy materials. In the event that you do not have a control number, please contact your broker, bank, or other nominee as soon as possible and no later than May 13, 2021,25, 2023, at least 30 minutes prior to the start of the meeting, so that you can be provided with a control number and gain access to the meeting. If you do not have a control number at the time of the meeting, you will be able to enter the meeting as a quest but not vote or ask questions.
Submitting Questions at the Virtual Annual Meeting. StockholdersThe Company will address questions submitted by stockholders during the 2023 Annual Meeting, which may submit questionsbe submitted in writing during the 2021 Annual Meeting onmeeting at www.virtualshareholdermeeting.com/BLI2021.CELL2023. Stockholders will need their unique control number which appears on theirthe Notice the proxy card (printed in the box and marked by the arrow) and the instructions that accompanied the proxy materials.
As part of theAt our 2023 Annual Meeting, we will not hold a short live question and answer session, during which we intend to answer questions submitted during the meeting in accordance with the Annual Meeting’s Rules of Conduct that are pertinent to the Company and the meeting matters, as time permits. Questions and answers will be grouped by topic and substantially similar questions will be grouped and answered once. In order to promote fairness, efficient use of the Company’s resources and in order to ensure all stockholders are responded to, we will respond to up to two (2) questions from a single stockholder.
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session.
The Annual Meeting’s Rules of Conduct will be posted on the InvestorInvestors Relations section of the Company'sCompany’s website approximately two weeks prior to the day of the 20212023 Annual Meeting.
Technical Assistance. Beginning 10 minutes prior to the start of and during the virtual 20212023 Annual Meeting, we will have support team ready to assist stockholders with any technical difficulties they may have accessing or hearing the virtual meeting.
If you encounter any difficulties accessing the virtual meeting during the check-in or meeting time, please call the technical support number that will be posted on the Virtual Shareholder Meeting log in page.



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Who pays for costs relating to the proxy materials and annual meeting of stockholders?
The costs of preparing, assembling and mailing the proxy materials, along with the cost of posting the proxy materials on a website, are to be borne by us. In addition to the use of mail, our directors, officers and employees may solicit proxies personally and by telephone, facsimile and other electronic means. They will receive no compensation in addition to their regular salaries. We may request banks, brokers and other custodians, nominees and fiduciaries to forward copies of the proxy materials to their principals and to request authority for the execution of proxies. We may reimburse these persons for their expenses in so doing. The costs of holding the 20212023 Annual Meeting will also be borne by us.
What does it mean if I received more than one Notice?
If you receive more than one Notice, your shares may be registered in more than one name or in different accounts. Please follow the voting instructions on each Notice to ensure that all of your shares are voted.
Is my vote confidential?
Proxy instructions, ballots and voting tabulations that identify individual stockholders are handled in a manner that protects your voting privacy. Your vote will not be disclosed either within Berkeley Lights,PhenomeX Inc. (“Company”) or to third parties, except as necessary to meet applicable legal requirements, to allow for the tabulation of votes and certification of the vote, or to facilitate a successful proxy solicitation.
I share an address with another stockholder, and we received only one paper copy of the proxy materials. How may I obtain an additional copy of the proxy materials?
We have adopted a procedure called “householding,” which has been approved by the SEC. Under this procedure, we will deliver only one copy of our Notice, of Internet Availability of Proxy Materials, and for those stockholders that receivedrequested a paper copy of our proxy materials in the mail, one copy of our annual report to stockholdersAnnual Report and this proxy statement,Proxy Statement, to multiple stockholders who share the same address (if they appear to be members of the same family) unless we have received contrary instructions from an affected stockholder. Stockholders who participate in householding will continue to receive separate proxy cards if they receivedrequested a paper copy of proxy materials in the mail. This procedure reduces our printing and mailing costs. Upon written or oral request, we will promptly deliver a separate copy of the proxy materials and annual reportAnnual Report to any stockholder at a shared address to which we delivered a single copy of any of these documents. To receive a separate copy, or, if you are receiving multiple copies, to request that we only send a single copy of next year’s proxy materials and annual report,Annual Report, you may contact us as follows:
Berkeley Lights,PhenomeX Inc.
Attn: General Counsel &Corporate Secretary
5858 Horton Street, Suite 320
Emeryville, CA 94608
(510) 858-2855
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Stockholders who hold shares in street name may contact their brokerage firm, bank, broker-dealer or other nominee to request information about householding.

What does it mean if I receive more than one Notice?
If you receive more than one Notice, your shares may be registered in more than one name or in different accounts. Please follow the voting instructions on each Notice to ensure that all of your shares are voted.
How can I find out the results of the voting at the 20212023 Annual Meeting?
Preliminary voting results will be announced at the 20212023 Annual Meeting. In addition, final voting results will be published in a current report on Form 8-K that we expect to file within four business days after the 20212023 Annual Meeting. If final voting results are not available to us at that time, we intend to file a Form 8-K to publish preliminary results and, within four business days after the final results are known to us, file an amendment to the Form 8-K to publish the final results.
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BOARD OF DIRECTORS AND CORPORATE GOVERNANCE
Director Independence
Our common stock is listed on the Nasdaq Global Select Market (“Nasdaq”). Under the rules of Nasdaq, independent directors must comprise a majority of a listed company’s board of directors within one year of the completion of an offering.directors. In addition, the rules of Nasdaq require that, subject to specified exceptions, each member of a listed company’s audit committee, compensation & leadership development committee and nominating and& corporate governance committeescommittee (if any) be independent. Audit committee members and compensation & leadership development committee members must also satisfy the independence criteria set forth in Rule 10A-3 and Rule 10C-1, respectively, under the Exchange Act.Act of 1934, as amended (“Exchange Act”). Under the rules of Nasdaq, a director will only qualify as an “independent director” only if, in the opinion of that company’s board of directors, that person does not have a relationship that would interfere with the exercise of independent judgment in carrying out the responsibilities of a director.
To be considered to be independent for purposes of Rule 10A-3 and under the rules of Nasdaq, a member of an audit committee of a listed company may not, other than in his or her capacity as a member of the audit committee, the board of directors or any other board committee: (1) accept, directly or indirectly, any consulting, advisory or other compensatory fee from the listed company or any of its subsidiaries or (2) be an affiliated person of the listed company or any of its subsidiaries.
To be considered independent for purposes of Rule 10C-1 and under the rules of Nasdaq, the board of directors must affirmatively determine that each member of the compensation & leadership development committee is independent, including a consideration of all factors specifically relevant to determining whether the director has a relationship to the company which is material to that director’s ability to be independent from management in connection with the duties of a compensation & leadership development committee member, including, but not limited to: (i) the source of compensation of such director, including any consulting, advisory or other compensatory fee paid by the company to such director and (ii) whether such director is affiliated with the company, a subsidiary of the company or an affiliate of a subsidiary of the company.
Our board of directors undertook a review of its composition, the composition of its committees and the independence of our directors and considered whether any director has a material relationship with us that could compromise his or her ability to exercise independent judgment in carrying out his or her responsibilities. Based upon information requested from and provided by each director concerning his or her background, employment and affiliations, including family relationships, our board of directors has determined that none of our continuing directors, other than Drs. Hobbs and Rothman,Dr. Kadia, has a relationship that would interfere with the exercise of independent judgment in carrying out the responsibilities of a director and that each of these directors is "independent" as that term is defined“independent” under the applicable rules and regulations of the listing requirements and rules of the Nasdaq Global Select Market.Nasdaq. Dr. HobbsKadia is not considered independent because he is an employee of Berkeley Lights. Dr. Rothman is not considered independent because he received compensation in excess of $120,000 during eachthe CEO of the preceding three years pursuant to our strategic/scientific advisor consulting agreement with him.Company.
In making these determinations, our board of directors considered the current and prior relationships that each non-employee director has with our companyCompany and all other facts and circumstances our board of directors deemed relevant in determining their independence, including the beneficial ownership of our capital stock by each non-employee director, and the transactions involving them described in the section titled “Certain Relationships and Related Party Transactions.” There are no family relationships among any of our executive officers or directors.
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Section 301.3 of the California Corporations Code
Because our principal executive office is located in California, we must comply with Section 301.3 of the California Corporations Code which provides that a publicly held corporation, as defined in Section 301.3, that has its principal executive offices in California must have at least one female director by the close of 2020, and may be required to have as many as three female directors by the close of 2021, depending on the authorized number of directors. Failure to comply with Section 301.3 can lead to the imposition of fines. As of December 31, 2020, our board of directors included two female directors, and therefore we were in compliance with Section 301.3 for 2020.
Board of Directors Leadership Structure
Our board of directors is currently chaired by Michael Marks.Greg Lucier. As a general policy, our board of directors believes that separation of the positions of chairmanchairperson of our board of directors and Chief Executive OfficerCEO reinforces the independence of our board of directors from management, creates an environment that encourages objective oversight of management’s performance and enhances the effectiveness of our board of directors as a whole. As such, Eric HobbsSiddhartha Kadia serves as our Chief Executive OfficerCEO while Michael MarksGreg Lucier serves as the chairmanchairperson of our board of directors but is not an officer. We currently expect and intend the positions of chairmanchairperson (or chairperson) of our board of directors and Chief Executive OfficerCEO to continue to be held by separate individuals in the future.

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Role of Board of Directors in Risk Oversight
Our board of directors has an active role, as a whole and also at the committee level, in overseeing the management of our risks. Our board of directors is responsible for general oversight of risks and regular review of information regarding our risks, including litigation risks and operational risks. The compensation & leadership development committee is responsible for overseeing the management of risks relating to our executive compensation plans and arrangements. The audit committee is responsible for overseeing the management of risks relating to accounting matters and financial reporting. The nominating and& corporate governance committee is responsible for overseeing the management of risks associated with the independence of our board of directors and potential conflicts of interest. Although each committee is responsible for evaluating certain risks and overseeing the management of such risks, our entire board of directors is regularly informed through discussions from committee members about such risks. Our board of directors believes its administration of its risk oversight function has not negatively affected the board of directors’ leadership structure.
Board of Directors and Committee Meetings
During 2020,2022, our board of directors held seveneight meetings (including regularly scheduled and special meetings) and acted by written consent on four occasions,, conducted periodic update calls, and each director attended at least 75% of the aggregate of (i) the total number of meetings of our board of directors held during the period for which he or she served as a director and (ii) the total number of meetings held by all committees of our board of directors on which he or she served during the periods that he or she served. It is the policy of our board of directors to regularly have separate meeting times for independent directors without management. Although we do not have a formal policy regarding attendance by members of our board of directors at annual meetings of stockholders, we encourage, but do not require, our directors to attend.

Board of Directors Committees
Our board of directors has threefour standing committees: the audit committee; the compensation & leadership development committee; the nominating & corporate governance committee and the nominating and corporate governancetransaction committee. Each committee is governed by a written charter approved by our board of directors, and written charters for the audit committee, the compensation & leadership development committee and the nominating & corporate governance committee, which satisfiessatisfy the applicable rules of the SEC and the listing standards of Nasdaq, and isare available on our website at https://investors.berkeleylights.com/investors.phenomex.com/ in the “Corporate Governance”“Governance” section of our Investors Relations webpage. We intend to post any amendments to the charters for each committee on the same website. We believe that the composition of these committees meets the applicable criteria for independence under, and the functioning of these committees complies with the applicable requirements of, the Sarbanes-Oxley Act of 2002 (“Sarbanes-Oxley Act”), the Nasdaq rules and SEC rules and regulations. We intend to continue to comply with Nasdaq requirements with
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respect to committee composition of independent directors. Each committee has the composition and responsibilities described below.

Audit Committee
Our audit committee consists of Ms. Nelson, Ms. BoyceMr. Lucier, Mr. Chiminski and Mr. Marks.Silvester. The chair of our audit committee is Ms. Nelson, whom ourNelson. Our board of directors has determined that each member of the audit committee is an “audit committee financial expert” as that term is defined under the SEC rules implementing Section 407 of the Sarbanes-Oxley Act, of 2002, and possesses financial sophistication, as defined under the Nasdaq listing standards of the Nasdaq Global Select Market.standards. Our board of directors has also determined that each member of our audit committee, can read and understand fundamental financial statements in accordance with applicable requirements, and that each member of our Audit Committeeaudit committee satisfy the independence standards for audit committee membership established by applicable rules of the SEC and the Nasdaq listing standards of the Nasdaq Global Select Market.standards. In arriving at these determinations, the board of directors has examined each audit committee member’s scope of experience and the nature of their experience in the corporate finance sector.professional experience. During 2020,2022, our audit committee held nine meetings.
seven meetings.


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The responsibilities of our audit committee include:
appointing, engaging, retaining, overseeing, approving the compensation of and assessing the independence of our independent registered public accounting firm;
overseeing the work of our independent registered public accounting firm, including through the receipt and consideration of reports from such firm;
reviewing and discussing with management and the independent registered public accounting firm our annual and quarterly financial statements and related disclosures;
monitoring our internal control over financial reporting, disclosure controls and procedures and code of business conduct and ethics;
discussing our risk management policies;
promoting compliance with applicable legal and regulatory requirements;
reviewing and approving or ratifying any related person transactions;transactions and our investment policy; and
preparing the audit committee report required by SEC rules.

Compensation & Leadership Development Committee
Our compensation & leadership development committee consists of Dr. Hopfield, Mr. Lucier, Ms. Nelson and Mr. Moritz and Ms. Nelson.Silvester. The chair of our compensation & leadership development committee is Mr. Lucier.Dr. Hopfield. Each of Dr. Hopfield, Mr. Lucier, Mr. Moritz and Ms. Nelson and Mr. Silvester is a non-employee director, as defined by Rule 16b-3 promulgated under the Exchange Act and meets the requirements for independence under the current Nasdaq listing standards and SEC rules and regulations. During 2020,2022, our compensation & leadership development committee held six meetings andeight meetings. In 2022, FW Cook acted by written consent once. The Compensation Committee has retained Radford, a part of Aon plc's Rewards Solution Practice (“Aon”) as itsan independent compensation consultant to help the Compensation Committee establish and implement its compensation philosophy, to evaluate compensation proposals recommended by management, and to provide advice and recommendations on competitive market practices and specific compensation decisions for executive officers. The Compensation Committee retains and does not delegate any of its exclusive power to determine all matters of executive compensation and benefits, although the CEO and the People Success department present compensation and benefit proposals to the Compensation Committee. Aon works directly with the Compensation Committee (and not on behalf of management) to assist the Compensation Committee in satisfying its responsibilities and will undertake no projects for management except at the request of the Compensation Committee chair and in the capacity of the Compensation Committee’s agent. Aon performs no other consulting or other services for Berkeley Lights' management. For additional description of the Compensation Committee’s processes and procedures for consideration and determination of executive officer compensation, see "Executive Compensation".& leadership development committee.
The responsibilities of our compensation & leadership development committee include:
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reviewing and approving, or recommending that our board of directors approve, the compensation of our chief executive officerCEO and our other executive officers;
reviewing and recommending to our board of directors the compensation of our directors;
selecting independent compensation consultants and advisors and assessing whether there are any conflicts of interest with any of the committeescommittee’s compensation advisors; and
reviewing and approving, or recommending that our board of directors approve, incentive compensation and equity plans.plans; and
preparing the compensation & leadership development committee report required by SEC rules.

Nominating and Corporate Governance Committee
Our nominating and& corporate governance committee consists of Mr. Chiminski, Dr. Khandros, Mr. LucierHopfield and Mr. Moritz.Ms. Nelson. The chair of our nominating and& corporate governance committee is Mr. Lucier.Chiminski. Each member of our nominating and& corporate governance committee is independent within the meaning of the applicable listing standards, is a non-employee director, and is free from any relationship that would interfere with the exercise of his or her independent judgment. During 2020,2022, our nominating and& corporate governance committee held twofive meetings.
The responsibilities of our nominating and& corporate governance committee include:

identifying individuals qualified to become board members;
recommending to our board the persons to be nominated for election as directors and to be appointed to each of the board’s committees;
reviewing and making recommendations to the board with respect to management succession planning;
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developing and recommending to the board corporate governance principles; andguidelines;
overseeing a periodic evaluation of the board.board; and
recommending to our board regarding governance matters, including, but not limited to, the Company’s certificate of incorporation, bylaws, and the charters of the Company’s other committees.
It is the policy of the nominating and& corporate governance committee of our board of directors to consider recommendations for candidates to our board of directors from stockholders. The nominating and& corporate governance committee will consider candidates recommended by stockholders in the same manner as candidates recommended to the nominating and& corporate governance committee from other sources. The nominating and& corporate governance committee may propose to our board of directors a candidate recommended or offered for nomination by a stockholder as a nominee for election to our board of directors. In the future, the nominating and& corporate governance committee may pay fees to third parties to assist in identifying or evaluating director candidates.
The nominating and& corporate governance committee seeks to identify as candidates for director persons with a reputation for and record of integrity and good business judgment. The nominating and& corporate governance committee considers the nature of the expertise and experience required for the performance of the duties of a director of the Company, and such matters as the candidate’s relevant business and industry experience, professional background, age, current employment, community service and other board service.
In addition, the nominating and& corporate governance committee shall take into account all factors it considers appropriate in recommending candidates for election to the board of directors, which may include:
The current size and composition of our board of directors and the needs of our board of directors and the respective committees of our board of directors.
Such factors as character, integrity, judgment, diversity of background and experience, independence, area of expertise, corporate experience, length of service, potential conflicts of interest, other commitments and the like. The nominating and& corporate governance committee evaluates these factors, among others, and does not assign any particular weighting or priority to any of these factors.
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Factors required to meet any applicable rules, regulations or laws regarding the composition of public company directors, including California Senate Bills Nos. 979 and 826, and proposed rules such asthe Nasdaq Proposed Rule 5605(f).; and
Other factors that the nominating and& corporate governance committee deems appropriate.
The nominating and& corporate governance committee requires the following minimum qualifications to be satisfied by any nominee for a position on our board of directors:
The highest personal and professional ethics and integrity.integrity;
Proven achievement and competence in the nominee’s field and the ability to exercise sound business judgment.judgment;
Skills that are complementary to those of the existing board of directors.directors;
The ability to assist and support management and make significant contributions to the Company’s success.success;
An understanding of the fiduciary responsibilities that is required of a member of our board of directors and the commitment of time and energy necessary to diligently carry out those responsibilities.responsibilities; and
If the nominating and& corporate governance committee determines that an additional or replacement director is required, the nominating and& corporate governance committee may take such measures that it considers appropriate in connection with its evaluation of a director candidate, including candidate interviews, inquiry of the person or persons making the recommendation or nomination, engagement of an outside search firm to gather additional information, or reliance on the knowledge of the members of the nominating and& corporate governance committee, our board of directors or management. 
While

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Transaction Committee
Our Transaction Committee consists of Mr. Chiminski, Dr. Hopfield, Mr. Lucier and Ms. Nelson. The chair of our Transaction Committee is Mr. Chiminski. The Transaction Committee has the Company does not have a formal policy on diversity for membersauthority of the boardBoard to consider and evaluate certain strategic investments and other possible transactions. During 2022, our transaction committee held eleven meetings.

The responsibilities of directors,our transaction committee include:
considering and evaluating all proposals that might be received by us in connection with potential strategic transactions, including, but not limited to, financings, mergers, acquisitions, divestitures, joint ventures, and other similar transactions;
participating in and directing the nominatingnegotiation of the material terms and corporate governance committee valuesconditions of any such transaction;
considering any strategic alternatives to any such potential strategic transaction;
recommending to the needBoard the advisability of entering into a definitive agreement (and any ancillary agreements relating thereto) with respect to any such strategic transaction, or the advisability of pursuing any strategic alternatives, in each case subject to applicable law; provided that the Board shall approve or submit any such strategic transaction or alternative for diversitythe approval of director skillsour stockholders; and viewpoints when considering new candidates.
performing any other activities or responsibilities incidental to the foregoing.
Stockholder Recommendations for Nominations to the Board of Directors
A stockholder that wants to recommend a candidate for election to the board of directors should direct the recommendation in writing by letter to Berkeley Lights,PhenomeX Inc., attention of the Corporate Secretary, at 5858 Horton Street, Suite 320, Emeryville, California 94608. The recommendation should include the candidate’s name, home and business contact information, detailed biographical data, relevant qualifications, a signed letter from the candidate confirming willingness to serve, information regarding any relationships between the candidate and Berkeley Lights, Inc.the Company and evidence of the recommending stockholder’s ownership of our stock. Such recommendations must also include a statement from the recommending stockholder in support of the candidate, particularly within the context of the criteria for board membership, as well as personal references.
A stockholder that instead desires to nominate a person directly for election to the board of directors at an annual meeting of the stockholders must meet the deadlines and other requirements set forth in Section 2.03 of our amended and restated bylaws and the rules and regulations of the SEC. Section 2.4 of our amended and restated bylaws requires that a stockholder who seeks to nominate a candidate for director must provide a written notice to the Corporate Secretary of the Company not later than the close of business on the ninetieth (90th) day and not earlier than the close of business on the one hundred and twentieth (120th) day prior to the first anniversary of the preceding year’s annual meeting; provided, however, that in the event that the date of the annual meeting is advanced by more than thirty (30) days, or delayed by more than sixty (60) days, from the anniversary date of the previous year’s meeting, or if no annual meeting was held in the preceding year, notice by the stockholder to be timely must be so delivered not earlier than the close of business on the one hundred and twentieth (120th) day prior to such annual meeting and not later than the close of business on the later of the ninetieth (90th) day prior to such annual meeting and the close of business on the tenth (10th) day following the day on which public announcement of the date of such meeting is first made. That notice must state the information required by Section 2.4 of our amended and restated bylaws, and otherwise must comply with applicable federal and state law. Public announcement of an adjournment or
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postponement of an annual meeting shall not commence a new time period (or extend any time period) for the giving of a stockholder’s notice. The Corporate Secretary of the Company will provide a copy of the bylaws upon request in writing from a stockholder. “Public announcement” shall mean disclosure (a) in a press release released by us, provided that such press release is released by us following our customary procedures, is reported by the Dow Jones News Service, Associated Press or comparable national news service, or is generally available on internetInternet news sites, or (b) in a document publicly filed by us with the SEC pursuant to Section 13, 14 or 15(d) of the Exchange Act and the rules and regulations promulgated thereunder.
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Stockholders also have the right under our bylaws to propose director candidates for consideration by the committee or our board of directors and also directly nominate director candidates, without any action or recommendation on the part of the nominating and& corporate governance committee or our board of directors, by following the procedures set forth in the section of this proxy statementProxy Statement titled “Stockholder Proposals“Proposals of Stockholders For 20222024 Annual Meeting.”
Compensation Committee Interlocks and Insider Participation
During 2020,2022, Dr. Hopfield, Dr. Kadia, Mr. Lucier Mr. Moritz and Ms. Nelson served on our compensation & leadership development committee. NoneAt the time of their respective service on the compensation & leadership development committee, none of the members of our compensation & leadership development committee is or has beenwas an officer or employee of our company. In February 2022, Dr. Kadia agreed to be considered for a role as the Company’s CEO, at which point Dr. Kadia recused himself from all compensation & leadership development committee discussions and meetings. Dr. Kadia formally stepped down from the committee prior to his appointment as CEO, which occurred on March 9, 2022. None of our executive officers currently serves, or in the past year has served, as a member of the board of directors or compensation & leadership development committee (or other board committee performing equivalent functions) of any entity that has one or more of its executive officers serving on our board of directors or compensation & leadership development committee.
Corporate Governance Guidelines and Code of Business Conduct and Ethics
Our board of directors has adopted corporate governance guidelines (“Corporate Governance Guidelines.Guidelines”). These guidelines address items such as the qualifications and responsibilities of our directors and director candidates and corporate governance policies and standards applicable to us in general. In addition, our board of directors has adopted a code of business conduct and ethics (“Code of Business Conduct and EthicsEthics”) that applies to all of our employees, officers and directors, including our Chief Executive Officer,CEO, Chief Financial Officer (“CFO”) and other executive and senior financial officers. The full text of our Corporate Governance Guidelines and our Code of Business Conduct and Ethics is posted on our website at https://investors.berkeleylights.com/investors.phenomex.com/ in the “Corporate Governance”“Governance” section of our Investors Relations webpage. We intend to post any amendments to our Corporate Governance Guidelines or Code of Business Conduct and Ethics, and any waivers of our Code of Business Conduct and Ethics for directors and executive officers, on the same website.
Communications with the Board of Directors
Our board of directors believes that management speaks for Berkeley Lights, Inc.the Company. Individual board members may, from time to time, communicate with various constituencies that are involved with the Company, but it is expected that board members would do this with knowledge of management and, in most instances, only at the request of management.
In cases where stockholders and other interested parties wish to communicate directly with our non-management directors, messages can be sent to our Corporate Secretary, at 5858 Horton Street, Suite 320, Emeryville, California 94608. Our Corporate Secretary monitors these communications and will provide a summary of all received messages to the board of directors at each regularly scheduled meeting of the board of directors. Our board of directors generally meets on a quarterly basis. Where the nature of a communication warrants, our Corporate Secretary may determine, in his or her judgment, to obtain the more immediate attention of the appropriate committee of the board or non-management director, of independent advisors or of Company management, as our Corporate Secretary considers appropriate.
Our Corporate Secretary may decide in the exercise of his or her judgment whether a response to any stockholder or interested party communication is necessary.
This procedure for stockholder and other interested party communications with the non-management directors is administered by the Company’s nominating and& corporate governance committee. This procedure does not apply to (a) communications to non-management directors from officers or directors of the Company who are stockholders,
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(b) stockholder proposals submitted pursuant to Rule 14a-8 under the Exchange Act or (c) communications to the audit committee pursuant to the Complaint Procedures for Accounting and Auditing Matters.
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Derivatives Trading, Hedging, and Pledging Policies
Our insider trading policy (“Insider Trading Compliance PolicyPolicy”) provides that no employee, officer, or director may acquire, sell, or trade in any interest or position relating to the future price of Company securities, such as a put option, a call option, or a short sale, or engage in hedging transactions (including “zero cost collars”). In addition, our Insider Trading Compliance Policy provides that no employee, officer, or director may pledge Company securities as collateral to secure loans. This prohibition means, among other things, that these individuals may not hold Company securities in a “margin” account, which would allow the individual to borrow against their holdings to buy securities.

Board Diversity and Composition
The following table provides our Board diversity statistics as self-disclosed by our directors as of April 14, 2023.

Board Diversity Matrix (As of April 14, 2023)
Board Size:
Total Number of Directors:9
Part I: Gender IdentityFemaleMaleNon-BinaryDid not Disclose Gender
Directors261
Part II: Demographic Background
African American or Black
Alaskan Native or Native American
Asian1
Hispanic or Latino
Native Hawaiian or Pacific Islander
White24
Two or More Races or Ethnicities
LGBTQ+
Did Not Disclose Demographic Background2



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DIRECTOR COMPENSATION

The following table provides information regarding compensation earned during the fiscal year ended December 31, 2022 by each non-employee director for his or her service on the board of directors and any committee(s). Our employee director, Siddhartha Kadia, Ph.D., receives no additional compensation for serving on our board of directors. The compensation paid to Dr. Kadia for serving as our Chief Executive Officer and for his service as a non-executive director during the fiscal year ended December 31, 2022 prior to his appointment as Chief Executive Officer is set forth in the Summary Compensation Table in the Executive Compensation section below.

NameFees Earned
or Paid in
Cash ($)(1)
Stock
Awards
($)(2)
Option
Awards
($)(3)
All other compensation ($) (4)Total
($)
James Rothman, Ph.D. (5)
40,000 89,999 90,010 125,000 345,009 
Sarah Boyce (5) (6)
25,000 — — — 25,000 
Elizabeth Nelson (5)
67,000 89,999 90,010 — 247,009 
Igor Khandros, Ph.D. (5)
45,000 89,999 90,010 — 225,009 
Michael Moritz (5)
46,750 89,999 90,010 — 226,759 
Gregory Lucier (5) (7)
95,250 184,829 90,010 — 370,089 
John Chiminski (5)
57,500 89,999 90,010 — 237,509 
Jessica Hopfield, Ph.D. (5)
53,000 89,999 90,010 — 233,009 
________________
(1)The fees paid to non-employee directors are described below under the heading “Non-Employee Director Compensation Program.”
(2)Amounts shown represent the grant date fair value of restricted stock units (“RSUs”) granted during the fiscal year 2022 as calculated in accordance with Financial Accounting Standards Board (“FASB”) Accounting Standard Codification (“ASC”) Topic 718. The grant date fair value for RSUs is measured based on the closing price of the Company’s common stock on the date of grant. These amounts do not correspond to the actual value that may be recognized by the director upon vesting of the applicable awards or sale of the underlying shares of stock. Each non-employee director received an annual director award of 19,565 RSUs pursuant to the terms of the Director Compensation Program described below.
(3)Amounts shown represent the grant date fair value of stock options granted during the fiscal year 2022 as calculated in accordance with FASB ASC Topic 718. The grant date fair value for stock options is estimated using the Black-Scholes option pricing model. For a discussion of the valuation assumptions used to determine the grant date fair value of the stock options granted to our non-employee directors in fiscal year 2022, see Note 13 to our Consolidated Financial Statements set forth in our Annual Report. These amounts do not correspond to the actual value that may be recognized by the director upon exercise of the applicable awards or sale of the underlying shares of stock. Each non-employee director received an annual director award of 32,258 stock options pursuant to the terms of the Director Compensation Program described below.
(4)Represents the amount that Dr. Rothman received as compensation associated with his participation on our Strategic Scientific Advisory Board during fiscal year 2022.
(5)The following table shows the aggregate numbers of option awards (exercisable and unexercisable) and unvested stock awards held as of December 31, 2022 by each non-employee director who was serving as of December 31, 2022.
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NameOptions OutstandingUnvested RSUs Outstanding
James Rothman, Ph.D.194,658 19,565 
Elizabeth Nelson139,658 19,565 
Igor Khandros, Ph.D.37,158 19,565 
Michael Moritz37,158 19,565 
Gregory Lucier162,158 23,731 
John Chiminski37,158 22,015 
Jessica Hopfield, Ph.D.48,765 26,013 

(6)Ms. Boyce did not stand for re-election at our 2022 annual meeting of stockholders.
(7)Mr. Lucier received an additional 16,666 RSUs on March 10, 2022 with a grant date fair value of $94,830. The RSUs vest in four equal quarterly installments beginning on May 20, 2022, subject to Mr. Lucier’s continued service.


Non-Employee Director Compensation Program
Prior to our initial public offering, which was completed in July 2020, we did not have a formal director compensation policy and did not pay our non-employee directors compensation for their services as a director, other than reimbursing them for reasonable, out-of-pocket travel expenses incurred while attending meetings of our board of directors or occasionally granting an equity award to certain of our non-employee directors upon their initial appointment to our board of directors. Following the completion of our initial public offering in July 2020, we adoptedWe maintain a non-employee director compensation program, (the “Program”), which was subsequentlymost recently amended in Februaryduring fiscal quarter ending December 31, 2021 (“Director Program”), pursuant to which each of our non-employee directors will receivereceives cash and equity compensation as follows, in addition to the reimbursement for travel and other necessary business expenses incurred in the performance of their services to us:us set forth below. The Director Program was developed with input from our independent compensation consultant, FW Cook, regarding practices and compensation levels at comparable companies. It is designed to attract, retain and reward our non-employee directors.
Each non-employee director will receivereceives an annual cash retainer in the amount of $40,000 per year.
Any non-executive chairperson will receivereceives an additional annual cash retainer in the amount of $30,000 per year (which was increased to $40,000 per year in February 2021).year.
The chairperson of the audit committee will receivereceives additional annual cash compensation in the amount of $20,000 per year for such chairperson’s service on the audit committee. Each non-chairperson member of the audit committee will receivereceives additional annual cash compensation in the amount of $10,000 per year for such member’s service on the audit committee.
The chairperson of the compensation & leadership development committee will receivereceives additional annual cash compensation in the amount of $12,000 per year (which was increased to $15,000 per year in February 2021) for such chairperson’s service on the compensation & leadership development committee. Each non-chairperson member of the compensation & leadership development committee will receivereceives additional annual cash compensation in the amount of $6,000 per year (which was increased to $7,000 per year in February 2021) for such member’s service on the compensation & leadership development committee.
The chairperson of the nominating and& corporate governance committee will receive additional annual cash compensation in the amount of $10,000 per year for such chairperson’s service on the nominating and& corporate governance committee. Each non-chairperson member of the nominating and& corporate governance committee will receive additional annual cash compensation in the amount of $5,000 per year for such member’s service on the nominating and& corporate governance committee.
Under the Program,Upon each non-employee director will automatically be granted (i) an option to purchase 80,000 shares of our common stock (which was decreased to 4,900 shares of our common stock in February 2021) upon the director’s initial appointment or election to our board of directors, the non-employee director will automatically be granted $360,000 worth of equity in the form of (i) an option to purchase shares of the Company’s common stock (“Initial Option”) with a grant date fair value of $180,000, and (ii) effective in February 2021, an award of 4,900 restricted stock units,RSUs (“Initial RSU Award”) with a grant date fair value of $180,000, and referred to, collectively, as the “Initial Grant”. The Initial Grant,Option vests as to 1/36th of the underlying shares
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on a monthly basis over three years, and for any non-employee director who has been serving on our boardthe Initial RSU Award vests as to one third (1/3) of directors, (iii) an option to purchase 40,000 shares of our common stock (which was decreased to 4,900 shares of our common stock in February 2021), and (iv) effective in February 2021, an award of 1,400 restricted stock units, automaticallythe RSUs on the first anniversary of the grant date ofand in equal quarterly installments for the following two years thereafter, subject to continued service through each annual stockholder’s meeting thereafter, referred to, collectively, as the Annual Grant.applicable vesting date. Non-employee directors elected for the first time to the board of directors at an
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annual stockholder'sstockholder’s meeting will only receive the Initial Grant.
Each non-employee director who has been serving on our board of directors for at least six months, will automatically be granted on the date of each annual award as defined above. The Initial Grant will vest, with respectstockholder’s meeting (i) an option to the option award, as to 1/36thpurchase shares of the underlying shares onCompany’s common stock (“Annual Option”) with a monthly basis over three years,grant date fair value of $90,000, and with respect to the(ii) an award of restricted stock units as to one third (1/3) of the restricted stock units on the first anniversary of the(“Annual RSU Award”) with a grant date fair value of $90,000, in each case pursuant to the 2020 Plan and vest thereafter in equal quarterly installments forreferred to, collectively, as the following two years, subject to continued service through each applicable vesting date.“Annual Grant.” The Annual Grant will vestvests on the earlier of the first anniversary of the date of grant or the date of the next annual stockholder’s meeting to the extent unvested as of such date, subject to continued service through each applicable vesting date.
The exercise price per share of director optionsthe Initial Options and the Annual Options is equal to the fair market value of a share of our common stock on the grant date, and all annualInitial Grants and initial awards willAnnual Grants vest in full upon the consummation of a Change in Control (as defined in the 2020 Plan)Director Program).

The 2020 Plan includes a maximum annual limit of $500,000 of cash compensation and equity awards that may be paid, issued or granted to a non-employee director in any calendar year (increased to $1,000,000 in the calendar year in which the non-employee director joins the board of directors as a non-employee director). For purposes of these limitations, the value of an equity award is based on its grant date fair value (determined as of the grant date in accordance with FASB Topic 718).

Upon joining the board of directors in April 2023, Mr. Silvester became eligible to receive the cash compensation under the Director Compensation Table
The following table provides information regarding compensation earned duringProgram described above, and in connection with his appointment, the fiscal year ended December 31, 2020board of directors approved that Mr. Silvester will receive an initial equity award at the 2023 Annual Meeting on terms approved by each non-employee director for his or her service on the board of directors and any committee(s). Our employee directors receive no additional compensation for serving on our board of directors. The compensation paid to Eric D. Hobbs, Ph.D. for serving as our Chief Executive Officer is set forth in the Summary Compensation Table in the Executive Compensation section below.
NameFees Earned
or Paid in
Cash ($)(1)
Stock Option
Awards
($)(2)
All other compensation ($) (3)Total
($)
James Rothman, Ph.D.20,000 782,111 156,250 958,361 
Sarah Boyce25,000 — — 25,000 
Elizabeth Nelson33,000 — — 33,000 
Igor Khandros, Ph.D.22,500 — — 22,500 
Michael Marks45,000 — — 45,000 
Michael Moritz25,500 — — 25,500 
Gregory Lucier (4)
31,000 1,283,482 — 1,314,482 
John Gunn (5)
— — — — 
Ming Wu, Ph.D. (6)
— — — — 
Makoto Shintani (7)
— — — — 
________________
(1)The fees paid to non-employee directors are describes below under the heading "Non-Employee Director Compensation Policy."
(2)Amounts shown represents the grant date fair value of options granted during the fiscal year 2020 as calculated in accordance with ASC Topic 718. These amounts do not correspond to the actual value that maybe recognized by the director upon exerciselieu of the applicable awards or sale of the underlying shares of stock. We did not grant stock options to any other non-employee director in the fiscal year 2020. As of December 31, 2020, the aggregate number of equity awards held by each of our non-employee directors was as follows: Dr. Rothman held options to purchase 157,500 shares of our common stock and 375,000 shares of our common stock acquired upon the exercise of his option awards, Ms. Boyce held options to purchase 100,000 shares of our common stock, Ms. Nelson held options to purchase 107,500 shares of our common stock and 50,000 shares of our common stock acquired upon the early exercise of her option award, Mr. Lucier held options to purchase 125,000 shares of our common stock, and Mr. Gunn held options to purchase 50,000 shares of our common stock. None of our other non-employee directors held any equity awards as of December 31, 2020.
(3)Represents the amount that Dr. Rothman received as compensation associated with his participation on our Strategic Scientific Advisory Board during 2020.
(4)Mr. Lucier joined our board of directors in June 2020.
(5)Mr. Gunn resigned from our board of directors in June 2020.
(6)Dr. Wu resigned from our board of directors in July 2020.
(7)Mr. Shintani resigned from our board of directors in July 2020.Initial Grant described above.
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MATTERS TO COME BEFORE THE 20212023 ANNUAL MEETING
PROPOSAL ONE:
Election of Directors
Board of Directors Composition
Our board of directors currently consists of eightnine members, with staggered three-year terms, pursuant to our amended and restated certificate of incorporation and amended and restated bylaws. Directors in Class IIII stand for election at the 20212023 Annual Meeting. The terms of office of directors in Class III and Class IIIII expire at our Annual Meetingsannual meetings of Stockholdersstockholders to be held in 20222024 and 2025, respectively.
James Rothman is not standing for re-election at the 2023 respectively. Annual Meeting and, therefore, will retire from the board of directors at the 2023 Annual Meeting. The board of directors thanks Mr. Rothman for his many contributions and his valuable insight.
At the recommendation of our nominating and& corporate governance committee, our board of directors proposes that each of the Class IIII nominees named below and currently serving as a director in Class I, be elected as a Class IIII director for a three-year term expiring at our 2024 Annual Meeting2026 annual meeting of Stockholders,stockholders, or until such director’s successor is duly elected and qualified, or until his earlier death, resignation, disqualification or removal.
NameAgePosition
Gregory LucierJohn Chiminski (1)
5659Director
Elizabeth NelsonPeter Silvester (2)
6054Director
________________
(1)Chairperson and memberMember of our compensationaudit committee, nominating & corporate governance committee and chairperson and member of our nominating and corporate governancetransaction committee.
(2)ChairpersonMember of our audit committee and compensation & leadership development committee.
Director Nominees
John Chiminski has served as a member of our board of directors since May 2021. Mr. Chiminski has served as the Executive Chair of Catalent Pharma Solutions since July 2022 and previously served as its Chief Executive Officer from March 2009 to July 2022. From 2007 to 2009, Mr. Chiminski was President and Chief Executive Officer of GE Medical Diagnostics (“GE”), a global business with sales of $1.9 billion. From 2005 to 2007, he served as Vice President and General Manager of GE Healthcare’s Global Magnetic Resonance Business, and from 2001 to 2005, as Vice President and General Manager of Global Healthcare Services. Earlier at GE, he held a series of cross-functional leadership positions in both manufacturing and engineering, including a GE Medical Systems assignment in France. Mr. Chiminski holds a B.S. from Michigan State University and an M.S. from Purdue University, both in electrical engineering, as well as a Master’s degree in Management from the Kellogg School of Management at Northwestern University. We believe Mr. Chiminski is qualified to serve on our board of directors due to his substantial expertise in advising and managing companies in various segments of the life science industry and his extensive experience as a business leader in the life science industry.

Peter Silvester has served as a member of our board of directors since April 2023. Mr. Silvester has most recently served as Senior Vice President and President of Life Sciences Solutions and Laboratory Products at Thermo Fisher Scientific Inc. (“Thermo Fisher”) from August 2017 to March 2022, and served as President of Biosciences at Thermo Fisher from January 2014 to August 2017. Previously, Mr. Silvester also held various positions at Life Technologies Corp. prior to its acquisition by Thermo Fisher, including Vice President of Global Enterprise Genomic Services and President of the Europe Middle East and Africa region. Mr. Silvester previously served on the board of directors of Coriell Life Sciences, Inc., a precision medicine company, and as an executive board member of Biocom California, an advocacy organization for the life sciences industry. Mr. Silvester received a B.S. in Applied Biology from the University of Hertfordshire. We believe Mr. Silvester is qualified to serve as a member
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of our board of directors because of his extensive executive and operational experience in the life sciences industry, including abroad and in emerging markets.

Continuing Directors
NameAgeClassTerm ExpirationPosition
Siddhartha Kadia, Ph.D. (1)
53I2024Director and CEO
Gregory Lucier (2)
58I2024Director
Elizabeth Nelson (3)
62I2024Director
Jessica Hopfield, Ph.D. (4)
58II2025Director
Igor Khandros, Ph.D.68II2025Director
Michael Moritz68II2025Director
________________
(1) Dr. Kadia served as a member of our audit committee and compensation & leadership development committee. In February 2022, Dr. Kadia agreed to be considered for a role as the Company’s CEO, at which point Dr. Kadia recused himself from all committee discussions and meetings. Dr. Kadia formally stepped down from both of the committees prior to his appointment as CEO, which occurred on March 9, 2022.
(2) Mr. Lucier is a member of our audit committee, compensation & leadership development committee and transaction committee.
(3) Ms. Nelson is a member of our audit committee, compensation & leadership development committee, nominating & corporate governance committee and transaction committee.
(4) Dr. Hopfield is a member of our compensation & leadership development committee, nominating & corporate governance committee and transaction committee.

Director NomineesSiddhartha Kadia, Ph.D. has served as a member of our board of directors since September 2021 and as the CEO of the Company since March 2022. Dr. Kadia was president and CEO of EAG Laboratories, a global scientific services company providing analytical testing and consulting solutions, which was acquired by Eurofins Scientific in December 2017. He continued to serve as the CEO of EAG laboratories until December 2018. Prior to EAG Laboratories, Dr. Kadia served in several senior roles including the president of Life Sciences Division at Life Technologies Corporation at the time of its acquisition by Thermo Fisher Scientific in February 2014. Prior to Life Technologies, Dr. Kadia was a management consultant at McKinsey & Company in the healthcare practice, assisting global medical device companies, and major healthcare providers. Dr. Kadia earned a B.E. in electronics and telecommunications from Gujarat University in India, an M.S. in biomedical engineering from Rutgers University, and a Ph.D. in biomedical engineering from Johns Hopkins University. He is currently a board member of the following publicly listed companies: ALS Limited (ASX:ALQ), Applied Technical Services (ATS) and NuVasive (NSDQ:NUVA). Dr. Kadia’s experience also includes prior public company board service as a director of Volcano Corporation (NSDQ: VOLC), Newport Corporation (NSDQ: NEWP), Horizon Discovery Group plc. (LSE: HZD), IsoPlexis Corporation (NSDQ: ISO) and Bioskryb Genomics, Inc. We believe Dr. Kadia is qualified to serve on our board of directors due to his significant strategic, financial, operational, and governance experience at other public and private life science and biotech companies.

Gregory Lucier has served as a member of our board of directors since June 2020. Mr. Lucier has served as the Chief Executive OfficerCEO of Corza Health, Inc., a privately-held company focused on acquiring companies and assets as part of a strategy to build a market-leading healthcare business, since April 2019. Prior to that, he served as Chief Executive Officer of NuVasive, Inc., a publicly-traded medical device company, from June 2015 to DecemberNovember 2018. Before joining NuVasive, Mr. Lucier was Chairman and Chief Executive OfficerCEO of Life Technologies Corporation (formerly Invitrogen Corporation), a publicly-traded global biotechnology company, from May 2003 until it was acquired by Thermo Fisher Scientific Inc. in February 2014. Prior to that, Mr. Lucier was a corporate officer at General Electric Company, where he served in a variety of leadership roles. Mr. Lucier isserved as Chairman of the boards of directors
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of Nuvasive,NuVasive, Inc., a publicly traded medical device company, from November 2018 to May 2021, and has served as a director of Catalent, Inc., a publicly-traded pharmaceutical company, since April 2015, and previously served on the boards of directors of Life Technologies Corporation from May 2003 to February 2014, of CareFusion Corporation, a publicly-traded medical device company, from August 2009 until its sale to Becton Dickinson in March 2015, and of Invuity, Inc. from October 2014 until its sale to Stryker in OctoberOctober 2018. On March 25, 2021, NuVasive announced that Mr. Lucier would be retiring from its Board of Directors effective May 18, 2021. Mr. Lucier received his B.S. in Industrial Engineering from Pennsylvania State University and his M.B.A. from Harvard Business School. We believe Mr. Lucier is qualified to serve on our board of directors due to his experience as an executive and director at medical device and life sciences companies.

Elizabeth Nelson has served as a member of our board of directors since September 2019. Since 2006, Ms. Nelson has served as principal of equusventures,EquusVentures, an investment fund that she founded. From 1996 until 2005, Ms. Nelson served as the Executive Vice President and Chief Financial Officer of Macromedia, Inc., software development company, where she also served as a director from January 2005 to December 2005. Ms. Nelson currently serves on the board of directors of Nokia Corporation, a Finnish publicly-traded telecommunications company, since July 2012, Upwork Inc., a publicly-traded technology company, since April 2015,, Virgin Group Acquisition Corp. II, a publicly traded investment company, since March 2021, and several private companies, including DAI Global, Noodle.ai and Smule.private companies. Previously, Ms. Nelson served on the board of directors of Nokia, a publicly traded telecommunications company, from May 2021 to April 2021, Zendesk, Inc., a publicly-traded software company, from July 2012 to May 2019, and Pandora Media, Inc., a music and software company, from
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July 2013 to June 2017. From 1996 until 2005, Ms. Nelson served as the Executive Vice President and CFO of Macromedia, Inc., software development company, where she also served as a director from January 2005 to December 2005. Ms. Nelson received her B.S. in Foreign Service from Georgetown University and her M.B.A. in Finance from the Wharton School at the University of Pennsylvania. In addition, Ms. Nelson completed the NACD certification in CyberSecurity Oversight offered by Carnegie Mellon University’s Software Engineering Institute in 2023. We believe that Ms. Nelson is qualified to serve on our board of directors due to her financial, accounting and operational experience from prior experience as an executive and director for various private and public technology companies.
Continuing Directors

NameAgeClassTerm ExpirationPosition
Sarah Boyce (1)
49II2022Director
Eric Hobbs44III2023Director and Chief Executive Officer
Igor Khandros (3)
66II2022Director
Michael Marks (1)
70III2023Director
Michael Moritz (2)(3)
66II2022Director
James Rothman70III2023Director
________________
(1) Member of our audit committee
(2) Member of our compensation committee
(3) Member of our nominating and corporate governance committee
Sarah BoyceJessica Hopfield, Ph.D. has served as a member of our board of directors since August 2019. Ms. BoyceDecember 2021. Dr. Hopfield currently serves as the president of J Hopfield Consulting, where since 2010 she has been advising technology firms that are working to commercialize their intellectual property and drive growth. From 1995 to 2009, Dr. Hopfield was a Partner of McKinsey & Company in its global pharmaceuticals and medical devices practice, where she served clients across pharmaceutical, biotechnology, medical device, and consumer industries. She also previously held management positions at Merck Sharp & Dohme Corp. in clinical development, outcomes research, and marketing. Dr. Hopfield earned a B.S. in Biology at Yale College, an MBA from the Harvard Graduate School of Business Administration and a Ph.D. in Neuroscience/Biochemistry in laboratory from the Rockefeller University. She is currently a Board member of Insulet Corporation (NASDAQ: PODD), Editas Medicine, Inc. (NASDAQ: EDIT), and Maravai LifeSciences (NASDAQ: MRVI). Dr. Hopfield’s experience includes prior public company board service as a director of Radius Health, Inc. (NASDAQ: RDUS). She also has served as President and Chief Executive OfficerChair of Avidity Biosciences LLC, a publicly-traded biotechnology company, since October 2019. Previously, Ms. Boyce served as the President and served onBoard of Trustees of the board of directors of Akcea Therapeutics, Inc., a publicly-traded biopharmaceutical company, from April 2018 to September 2019. Before that, Ms. Boyce was the Chief Business Officer of Ionis Pharmaceuticals, Inc., a publicly-traded biotechnology company, from January 2015 to April 2018. Prior to joining Ionis, Ms. Boyce was Vice President, Head of International Business Strategy and Operations at Forest Laboratories, Inc., a publicly-traded pharmaceutical company that was acquired by Actavis Pharma, Inc. from 2012 to 2014. She was Vice President, Global Head Nephrology Therapeutics Area of Alexion Pharmaceuticals Inc., a publicly-traded pharmaceutical company, from 2010 to 2011. She held various positions at Novartis Group AG, a publicly-traded Swiss healthcare company, including Vice President, Global Program Head, Pediatric and Specialty from 2000 to 2010. Ms. Boyce currently serves on the board of directors of Avidity Biosciences LLC, a privately-held biotechnology company and Ligand Pharmaceuticals Incorporated, a publicly-traded biopharmaceutical company. Ms. Boyce received her B.S. in Microbiology from the University of Manchester, England.Joslin Diabetes Center. We believe Ms. BoyceDr. Hopfield is qualified to serve on our board of directors due to her more than two decades of executive healthcare experience along with extensive experience as an executivepublic company independent director board expertise across a range of governance roles and director of public and private companies in the biotechnology industry.committees.
Eric D. Hobbs, Ph.D. has served as a member of our board of directors and as our Chief Executive Officer since March 2017. Previously, Dr. Hobbs served as our Senior Vice President, Operations and Consumables, from March 2016 to March 2017, as our Vice President, Operations and Consumables, from February 2015 to March 2016, and as Senior Director, Research and Development, from May 2013 until February 2015. Prior to Berkeley Lights, Dr. Hobbs held roles of increasing responsibility at FormFactor, Inc., a publicly-traded semiconductor technology company, serving most recently as Senior Director of Product Management. Dr. Hobbs received his B.A. in Liberal Arts, with minors in Mathematics and Physics from Saint Mary’s College of California, his B.S. in Mechanical Engineering from the University of Southern California, and his M.S. and Ph.D. in Mechanical Engineering, Microelectromechanical Systems, from the University of California, Berkeley. We believe Dr. Hobbs is qualified to serve on our board of directors due to his experience as our Chief Executive Officer and his educational background.
Igor Khandros, Ph.D. has served as a member of our board of directors since 2011. Dr. Khandros has served as Chief Executive OfficerCEO of Nutcracker Therapeutics, Inc., a privately-held biotechnology company, since October 2017. Dr. Khandros is one of our Co-Founders and, from 2011 to March 2017, served as our Chief Executive Officer.CEO. Previously, Dr. Khandros founded and served on the boards of directors of two publicly-traded companies,
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FormFactor, Inc., a publicly-traded semiconductor technology company, and Tessera Technologies, Inc., a technology licensing company. Dr. Khandros received an M.S.-equivalent degree in Metallurgical Engineering from Kiev Polytechnic Institute in Kiev, Ukraine, and a Ph.D. in Metallurgy from Stevens Institute of Technology. We believe Dr. Khandros is qualified to serve on our board of directors due to his educational background, his experience as a founder and former executive of our Company and his extensive experience as a founder, executive and board member at other public and private technology companies.
Michael Marks has served as a member of our board of directors since April 2014. Mr. Marks has served as Founding Managing Partner of WRVI Capital, a technology venture capital fund, since 2014, as Founding Managing Partner of WRV Capital, a technology venture capital fund, since 2014 and as Chief Executive Officer, Chairman and Founder of Katerra, Inc., or Katerra, a technology company, from March 2015 to May 2020. Previously, Mr. Marks served as a founding partner of Paxion Capital, a private partnership, from March 2015 until March 2020, was a Founding Partner of Riverwood Capital, a private equity firm, from March 2007 until May 2019 and a Partner and Senior Advisor at Kohlberg Kravis Roberts & Co., a private equity firm, from 2006 to 2007. Prior to these roles, Mr. Marks served as Chief Executive Officer of Flextronics from 1994 until 2006 and as member of its board of directors from 1994 to 2007. Mr. Marks currently serves on the board of directors of several companies, including Bossa Nova, a privately-held robotics company, The Melt, a restaurant chain, Finicast, a financial planning and analysis software solution company, and Smart Frog/Canary, a smart home security company. Mr. Marks previously served on the board of directors of publicly-traded companies, Schlumberger Limited, an oilfield services company, from January 2015 to July 2019, GoPro, Inc., a technology company from February 2011 to April 2017 and SanDisk Corporation, a manufacturer of flash memory products from 2003 to September 2016. Mr. Marks received his B.A. and M.A. in Psychology from Oberlin College and his M.B.A. from Harvard Business School. We believe Mr. Marks is qualified to serve on our board of directors due to his extensive experience as chief executive officer and member of the boards of directors of numerous companies, his expertise in financial and accounting matters and his investment experience.
Michael Moritz has served as a member of our board of directors since April 2015. Mr. Moritz has been a Managing Partner of Sequoia Capital, a venture capital fund, since 1986. Mr. Moritz currently serves on the board of directors
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of numerous private companies, including Stripe, Inc., a financial technology company, Instacart, a grocery delivery technology company, Charlotte Tilbury, a cosmetics company, GameFly, a technology company,and Klarna, a financial technology company, and Group Nine Media, a digital media company. Previously, Mr. Moritz served on the board of directors of LinkedIn Corporation, a publicly-traded professional networking company, from 2011 to June 2016 and Green Dot Corporation, a publicly-traded financial technology company, from 2003 to May 2016. Mr. Moritz received his M.A. in History from Christ Church, Oxford. We believe Mr. Moritz is qualified to serve on our board of directors due to his background in and over 25 years’ experience with the venture capital industry, providing guidance and counsel to a range of internetInternet and technology companies, and service on the boards of directors of various private and public companies.

James Rothman, Ph.D.
has served as a member of our board of directors since May 2016. Dr. Rothman has been a faculty member at Yale University since 2008, where he serves as the Sterling Professor of Cell Biology, Chairman of the Yale School of Medicine’s Department of Cell Biology and is the Director and founder of the Nanobiology Institute. Dr. Rothman served as Chief Scientist of GE Healthcare from 2002-2006 and as advisor to GE Research until 2013. Previously, Dr. Rothman founded and chaired the Department of Cellular Biochemistry and Biophysics at Memorial Sloan-Kettering Cancer Center from 1991 until 2004, where he held the Paul A. Marks Chair and served as Vice-Chairman. Previously, Dr. Rothman was the Wu Professor of Chemical Biology in the Department of Physiology and Cellular Biophysics at Columbia University and Director of Columbia University’s Sulzberger Genome Center, from 2004 to 2008. Dr. Rothman was awarded the 2013 Nobel Prize in Physiology or Medicine, for his work related to cell biology. Dr. Rothman currently serves on the boards of directors for various private biotechnology companies. He chairs the healthcare advisory board of Arsenal Capital Partners, a private equity fund, and previously served as an advisor to R&D leadership at GSK and Johnson & Johnson, among other large pharma companies. Dr. Rothman received his B.A. in Physics from Yale College and his Ph.D. in Biochemistry from Harvard University. We believe that Dr. Rothman is qualified to serve on our board of directors due to his educational background and extensive experience in biochemistry and cell biology, as well as his experience as an executive of healthcare and biotechnology companies.
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Vote Required

The election of Class IIII directors requires a plurality of the votes cast to be approved. "Withhold”“Withhold” votes and broker non-votes are not considered votes cast for the foregoing purpose, and will have no effect on the election of the nominees.
RECOMMENDATION OF THE BOARD OF DIRECTORS
THE BOARD OF DIRECTORS UNANIMOUSLY
RECOMMENDS A VOTE FOR THE ELECTION OF EACH NOMINEE UNDER PROPOSAL ONE

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PROPOSAL TWO:
Ratification of Independent Registered Public Accounting Firm
Our audit committee has appointed KPMG LLP, as our independent registered public accounting firm to audit our financial statements for our fiscal year ending December 31, 2021.2023. KPMG LLP has served as our independent registered public accounting firm since 2015.
Stockholder ratification of the appointment of KPMG LLP is not required by our bylaws or other applicable legal requirements. However, our board of directors is submitting the appointment of KPMG LLP to our stockholders for ratification as a matter of good corporate governance. In the event that this appointment is not ratified by the affirmative vote of the holders of a majority in voting power of the votes cast (meaning the number of shares voted “for” a proposal must exceed the number of shares voted “against” such proposal), such appointment will be reconsidered by our audit committee and the audit committee may reconsider whether it should appoint another independent registered public accounting firm. Even if the appointment is ratified, our audit committee, in its sole discretion, may appoint another independent registered public accounting firm at any time during our fiscal year ending December 31, 20212023 if our audit committee believes that such a change would be in the best interests of the Company and its stockholders.
We expect that representatives of KPMG LLP will attend the 20212023 Annual Meeting and will have the opportunity to make a statement if they so desire and to respond to appropriate questions.
Fees Paid to the Independent Registered Public Accounting Firm
The following table presents fees for professional audit services and other services rendered to us by KPMG LLP for our fiscal years ended December 31, 20192021 and 2020.2022:
20192020 20212022
Audit Fees (1)
Audit Fees (1)
$355,000 $1,870,000 
Audit Fees (1)
$1,847,500 $1,155,000 
Audit-Related Fees (2)
Audit-Related Fees (2)
— — 
Audit-Related Fees (2)
— — 
Tax Fees (3)
Tax Fees (3)
71,248 57,945 
Tax Fees (3)
82,635 107,631 
All Other Fees (4)
All Other Fees (4)
— 1,780 
All Other Fees (4)
1,780 — 
TotalTotal$426,248 $1,929,725 Total$1,931,915 $1,262,631 
________________
(1)Audit Fees include billed and unbilled fees forrelate to professional services rendered in connection with the audit of our annual consolidated financial statements included in our annual reportAnnual Reports on Form 10-K, and registration statement on Form S-1, the review of the unaudited interim consolidated financial statements included in our quarterly reportQuarterly Reports on Form 10-Q, and other professional services related to our initial public offering and various accounting consultations.provided in connection with regulatory filings, including registration statements filed with the SEC.
(2)Audit-Related Fees consist of fees for assurance and related services that are reasonably related to the performance of the audit or review of our financial statements and are not reported under Audit Fees. There were no Audit-Related Fees incurred in 20192021 or 2020.2022.
(3)Tax Fees include fees in connection with tax consulting and compliance services.
(4)All Other Fees relate to other products and professional services provided by KPMG LLP and include an annual subscription to an online technical accounting research tool incurred in 2020.tool.





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Auditor Independence
In 2020,2022, there were no other professional services provided by KPMG LLP that would have required our audit committee to consider their compatibility with maintaining the independence of KPMG LLP.
Audit Committee Policy on Pre-Approval of Audit and Permissible Non-Audit Services of Independent Registered Public Accounting Firm
Our audit committee has established a policy governing our use of the services of our independent registered public accounting firm. Under the policy, our audit committee is required to pre-approve all audit and permissible non-audit services performed by our independent registered public accounting firm in order to ensure that the provision of such services does not impair such accounting firm’s independence. All fees paid to KPMG LLP for our fiscal years ended December 31, 20202022 and 20192021 were pre-approved by our audit committee.

Vote Required
The ratification of the appointment of KPMG LLP requires the affirmative vote of the holders of a majority in voting power of the votes cast (meaning the number of shares voted “for” athe proposal must exceed the number of shares voted “against” such proposal). Abstentions and broker non-votes are not considered votes cast for the foregoing purpose, and will have no effect on the vote for this proposal.

RECOMMENDATION OF THE BOARD OF DIRECTORS
THE BOARD OF DIRECTORS UNANIMOUSLY
RECOMMENDS A VOTE FOR THE RATIFICATION OF KPMG LLP
AS THE COMPANY’S INDEPENDENT PUBLIC ACCOUNTING FIRM UNDER PROPOSAL TWO
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AUDIT COMMITTEE REPORT

Pursuant to rules adopted by the Securities and Exchange Commission (the “SEC”)SEC designed to improve disclosures related to the functioning of corporate audit committees and to enhance the reliability and credibility of financial statements of public companies, the audit committee of our board of directors submits the report below. The material in this report is not “soliciting material,” is not be deemed “filed” with the SEC, and is not to be incorporated by reference into any filing made by Berkeley Lights, Inc. (the “Company”)the Company under the Securities Act of 1933, as amended, or the Securities Exchange Act of 1934, as amended, whether made before or after the date hereof and irrespective of any general incorporation language in any such filing, except to the extent that the Company specifically incorporates it by reference in such filing.
Audit Committee Report to Stockholders
The audit committee of the board of directors is responsible for providing independent, objective oversight of the Company’s accounting functions and internal controls. The audit committee is composed of threefour directors, each of whom is independent as defined under the rules of the Nasdaq Global Select Market. Our board of directors has determined that each member of the audit committee is independent and that Ms. Nelson qualifies as an “audit committee financial expert” under the SEC rules. The audit committee operates under a written charter approved by the board of directors and held sevennine meetings in fiscal year 2020.2022. A copy of the charter is available on the Company’s website at https://investors.berkeleylights.com/investors.phenomex.com/ in the “Corporate Governance”“Governance” section of the Investors Relations webpage.
Management is responsible for the Company’s internal controls over financial reporting, disclosure controls and procedures and the financial reporting process. KPMG LLP is responsible for performing an independent audit of the Company’s financial statements in accordance with Public Company Accounting Oversight Board (“PCAOB”) standards and to issue reports thereon. The audit committee’s responsibility is to monitor and oversee these processes. The audit committee has established a mechanism to receive, retain and process complaints on auditing,
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accounting and internal control issues, including the confidential, anonymous submission by employees, vendors, customers and others of concerns on questionable accounting and auditing matters.
In connection with these responsibilities, the audit committee met with management and KPMG LLP to review and discuss the December 31, 20202022 audited financial statements. The audit committee also discussed with KPMG LLP the matters required to be discussed by the applicable requirements of the PCAOB and the SEC. In addition, the audit committee received the written disclosures from KPMG LLP required by applicable requirements of the PCAOB regarding KPMG LLP’s communications with the audit committee concerning independence, and the audit committee has discussed with KPMG LLP its independence from the Company and its management.
Based on the audit committee’s discussions with management and KPMG LLP, and the audit committee’s review of the representations of management and KPMG LLP, the audit committee recommended that the board of directors include the audited financial statements in the Company’s Annual Report on Form 10-K for the fiscal year 20202022 filed with the SEC.
The audit committee also has appointed, subject to stockholder ratification, KPMG LLP as the Company’s independent registered public accounting firm for the fiscal year 2021.2023.
Respectfully submitted,
THE AUDIT COMMITTEE OF
THE BOARD OF DIRECTORS OF BERKELEY LIGHTS,PhenomeX INC.
Elizabeth Nelson, Chairperson
Sarah BoyceGreg Lucier
Michael MarksJohn Chiminski
Peter Silvester
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Table
PROPOSAL THREE:
Advisory Vote on the Compensation of contentsNamed Executive Officers

As part of the commitment of our board of directors to excellence in governance, and as required by Section 14A of the Exchange Act, our board of directors is providing the stockholders with an opportunity to approve, on a non-binding, advisory basis, the compensation of our named executive officers

As more fully described in the section of this Proxy Statement titled “Executive Compensation,” including related compensation tables, our executive compensation program is designed to attract, retain and motivate talented individuals with the executive experience and leadership skills necessary for us to increase stockholder value.

The following proposal, commonly known as a “say on pay” proposal, gives our stockholders the opportunity to vote to approve, on a non-binding, advisory basis, the compensation of our named executive officers. This vote is not intended to address any specific item of compensation or the compensation of any particular officer, but rather the overall compensation of our named executive officers and our compensation philosophy, policies and practices, as discussed in this Proxy Statement. Accordingly, we are asking our stockholders to vote for the following resolution:

“RESOLVED, that the company’s stockholders approve, on a non-binding, advisory basis the compensation of the company’s named executive officers, as disclosed in this Proxy Statement, including the Executive Compensation section, compensation tables and narrative discussion.”

Before you vote, we recommend that you read the Executive Compensation section of this Proxy Statement for additional details on our executive compensation programs and philosophy.

This vote is advisory, and therefore not binding on the Company, our board of directors or the compensation & leadership development committee. However, our board of directors and compensation & leadership development committee value the opinions of our stockholders and intend to take into account the outcome of the vote when considering future compensation decisions for our named executive officers.

Vote Required
Approval of the compensation of our named executive officers on a non-binding, advisory basis requires the affirmative vote of the holders of a majority in voting power of the votes cast. Broker non-votes and abstentions will have no effect on this proposal.


RECOMMENDATION OF THE BOARD OF DIRECTORS
THE BOARD OF DIRECTORS RECOMMENDS
A VOTE FOR APPROVAL ON A NON-BINDING, ADVISORY BASIS, OF THE
COMPENSATION OF THE COMPANY’S NAMED EXECUTIVE OFFICERS UNDER PROPOSAL THREE
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EXECUTIVE OFFICERS
The following table sets forth certain information about our executive officers and their respective ages as of March 12, 2021.30, 2023. Officers are elected by the board of directors to hold office until their successors are elected and qualified.
NameAgePosition
Eric D. Hobbs,
Siddhartha Kadia, Ph.D.(1)
4453Chief Executive OfficerCEO and Director
Shaun M. HoltMehul Joshi (1)(2)
4362CFO
Scott Chaplin (3)
56Chief FinancialLegal Officer
Keith J. Breinlinger,
Eric D. Hobbs, Ph.D.(4)
46Chief Technology Officer
Stuart L. Merkadeau59General Counsel
Matthew W. Rosinack48SeniorExecutive Vice President Chief Accounting Officerof Integrations and Business Transformation
________________
(1) Dr. Kadia was appointed CEO on March 9, 2022.
(2) Mr. Holt advisedJoshi was appointed CFO effective July 25, 2022.
(3) Mr. Chaplin was appointed Chief Legal Officer on September 19, 2022.
(4) Dr. Hobbs was appointed Executive Vice President of Integrations and Business Transformation on April 1, 2023.

Siddhartha Kadia, Ph.D. has served as a member of our board of directors since September 2021 and as the CEO of the Company on February 2, 2021since March 2022. Dr. Kadia was president and CEO of his intentEAG Laboratories, a global scientific services company providing analytical testing and consulting solutions, which was acquired by Eurofins Scientific in December 2017. He continued to resignserve as Chief Financial Officer upon the Company’s filingCEO of EAG laboratories until December 2018. Prior to EAG Laboratories, Dr. Kadia served in several senior roles including the president of Life Sciences Division at Life Technologies Corporation at the time of its Annual Report on Form 10-K for its fiscal 2020.Kurt Woodacquisition by Thermo Fisher Scientific in February 2014. Prior to Life Technologies, Dr. Kadia was appointeda management consultant at McKinsey & Company in the healthcare practice, assisting global medical device companies, and major healthcare providers. Dr. Kadia earned a B.E. in electronics and telecommunications from Gujarat University in India, an M.S. in biomedical engineering from Rutgers University, and a Ph.D. in biomedical engineering from Johns Hopkins University. He is currently a board member of the following publicly listed companies: ALS Limited (ASX:ALQ), Applied Technical Services (ATS) and NuVasive (NSDQ:NUVA). Dr. Kadia’s experience also includes prior public company board service as Chief Financial Officer effective March 15, 2021.a director of Volcano Corporation (NSDQ: VOLC), Newport Corporation (NSDQ: NEWP), Horizon Discovery Group plc. (LSE: HZD), IsoPlexis Corporation (NSDQ: ISO) and Bioskryb Genomics, Inc.

For the biography of Eric D. Hobbs, Ph.D., see “Matters to Come Before the 2021 Annual Meeting—Proposal One—Election of Directors—Continuing Directors.”
Shaun M. HoltMehul Joshi has served as our Chief Financial Officer since March 2016.July 25, 2022. Mr. Joshiis an experienced finance executive with a strong background in the life sciences and technology industries, including at companies with a combination of equipment, software, and services business units. Previously, Mr. Holt served as our Vice President, Head of Finance and Accounting from November 2015 to March 2016. Mr. Holt previouslyhe served in a varietynumber of senior finance leadershipand accounting roles since May 1, 2011 at Illumina, Inc.,ResMed Incorporated, a publicly traded life sciences analyticsdigital health and cloud-connected medical device company, from July 2008 until November 2015, including Senior Director,most recently as senior vice president, Finance & Head of Finance, Global Business Units and Research and Development, Finance Director, Head of EMEA FinanceFP&A & Accounting Services. At ResMed, Mr. Joshi directed global finance teams that played an integral role in driving transformational growth supporting multi-channel business models including pricing and Headprofitability, specifically for their lease-to-own CPAP market, M&A, Operations, IR & Treasury, and was interim CFO of Commercial Financial Planningthe Software as a Services (SaaS) business line. Prior to ResMed, Mr. Joshi served in leadership positions of increasing responsibility at Gilead Sciences where he supported the exponential growth of the company, evaluated, and Analysis.integrated strategic acquisitions, and implemented tools, processes, and analytics across the company. Mr. Holt also previously served as Manager of Financial PlanningJoshi earned his B.A. in Accounting from Rutgers University, and Analysis at Websense, Inc. from June 2005 until July 2008. Mr. Holt was a Senior Financial Analyst at Kyocera Communications, Inc. from 2004 to 2005, and Cost Accountant, Cost Analyst and Financial Analyst at Macromedia, Inc. from 2001 to 2004. Mr. Holt received his B.B.A.MBA in Finance and Management from California State University, Stanislaus.Rutgers University.

Keith J. Breinlinger, Ph.D.Scott Chaplin has served as our Chief TechnologyLegal Officer since March 2013.September 19, 2022. Previously, Dr. BreinlingerMr. Chaplin served as Chief Legal and People Officer at Shield AI, a technology company focused on AI software, robotics, and autonomy solutions for the national security sector. Under his leadership, the company scaled organically and through strategic
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acquisitions from a startup company to more than $2B in valuation. Prior to Shield AI, Mr. Chaplin was chief legal and human resources officer at Vista Outdoor, Inc., where he led numerous acquisitions leading to a Productdiverse portfolio of 30 different company brands. In addition, Mr. Chaplin served as the Chief Legal Officer for Alliant Techsystems and System ArchitectStanley, Inc. He also worked as an adjunct professor of law at FormFactorAmerican University, Washington College of Law and as an attorney for Morgan, Lewis & Bockius LLP and Reed Smith LLP, both in Washington, D.C. Mr. Chaplin earned a B.A. degree from July 2005 until March 2013,the University of Massachusetts Amherst and a Mechanical Architect at TeradyneJuris Doctor degree from 1996 to 2005. From 1993 until 1995, Dr. Breinlinger was a Mechanical Designer at iRobot. Dr. Breinlinger received his B.S., M.S. and Ph.D. in Mechanical Engineering from the Massachusetts InstituteAmerican University, Washington College of Technology.Law.

Stuart L. MerkadeauEric D. Hobbs, Ph.D. has served as our General CounselExecutive Vice President of Integrations and Business Transformation since June 2015. Mr. Merkadeau previouslyApril 1, 2023. Previously, Dr. Hobbs served as our President of Global Operations from March 2022 to March 2023, Chief Executive Officer from March 2017 to March 2022, Senior Vice President, General CounselOperations and Corporate Secretary of FormFactor from October 2002 until May 2015, and as Vice President, Intellectual Property of FormFactor from July 2000 to September 2002. Prior to this, Mr. Merkadeau was a partner at Graham & James LLP, a private law firm, where he oversaw a variety of licensing, corporate, and litigation matters across a broad range of technologies. Mr. Merkadeau received his B.S. in Industrial Engineering from Northwestern University and his J.D. from the University of California, Los Angeles School of Law. He is admitted to practice law in California and is registered before the U.S. Patent and Trademark Office.

Matthew W. Rosinack has served as our Senior Vice President and Chief Accounting Officer since September 2020. Mr. Rosinack previously served as the Company’s Vice President, Finance and Accounting from February 2018 until September 2020, and as the Company’s Corporate ControllerConsumables, from March 2016 untilto March 2017, as our Vice President, Operations and Consumables, from February 2018. Prior to Berkeley Lights, from October 20122015 to March 2016, Mr. Rosinack held roles of increasing responsibility at ZELTIQ Aesthetics, Inc., includingand as Vice PresidentSenior Director, Research and Corporate Controller. Mr. Rosinack alsoDevelopment, from May 2013 until February 2015. Prior to PhenomeX, Dr. Hobbs held roles of increasing responsibility at FormFactor, Inc., from April 2011 to October 2012, including Corporate Controller anda publicly-traded semiconductor technology company, serving most recently as Senior Director
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Corporate Accounting. In addition, Mr. Rosinack spent more than 14 years in public accounting practice with PricewaterhouseCoopers LLP. Mr. RosinackProduct Management. Dr. Hobbs received his B.A. in AccountingLiberal Arts, with minors in Mathematics and Physics from Ohio Wesleyan University.Saint Mary’s College of California, his B.S. in Mechanical Engineering from the University of Southern California, and his M.S. and Ph.D. in Mechanical Engineering, Microelectromechanical Systems, from the University of California, Berkeley.




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EXECUTIVE COMPENSATION

As an “emerging growth company”a smaller reporting company, we are not required to make certain disclosures related to executive compensation, such as defined in the JOBS Act,providing a Compensation Discussion and Analysis, however we have optedelected to complyprovide our stockholders with thea description of our executive compensation rules applicable to “smaller reporting companies,” as such term is defined under the under the Securities Act of 1933, as amended, (the “Securities Act”), which require compensation disclosure for our principal executive officerprograms, and the two most highly compensated executive officers other thandecisions made by our principal executive officer. The following Summary Compensation Table summarizescompensation & leadership development committee (which we refer to as the total compensation of“Compensation Committee” in this “Executive Compensation” section) under those programs for our named executive officers (“NEOs”) for all services rendered to the Company in all capacities for the fiscal years ending December 31, 20202022, including executive compensation tables and 2019. As an “emerging growth company,” we are not required to include a Compensation Discussion and Analysis section and have elected to comply with the scaled disclosure requirements applicable to emerging growth companies.other information.

Executive Summary

Our Executives
Our NEOs for the fiscal year 2020 were as follows:2022 were:
Siddhartha Kadia, Ph.D., our Chief Executive Officer;
Eric D. Hobbs, Ph.D., our former Chief Executive Officer;Officer and current Executive Vice President of Integrations and Business Transformation;
Keith Breinlinger, Ph.D.,Mehul Joshi, our Chief Technology Officer;Financial Officer (“CFO”); and
Matthew W. Rosinack,Scott Chaplin, our Chief AccountingLegal Officer. and Corporate Secretary.
Summary Compensation Table
Name and Principal PositionYearSalary
($)(1)
Bonus
($)(2)
Stock
Option
Awards
($)(3)
Non-equity
Incentive Plan
Compensation
($)
Total
($)
Eric D. Hobbs, Ph.D.2020540,025 — 480,579 — 1,020,604 
Chief Executive Officer2019513,750 — 6,934,085 — 7,447,835 
Keith J. Breinlinger, Ph.D.2020395,904 — 238,151 — 634,055 
Chief Technology Officer2019375,000 — 715,694 64,453 1,155,147 
Matthew W. Rosinack2020286,952 24,400 1,113,374 — 1,424,726 
Chief Accounting Officer
________________Management Transition
(1)2022 was a year of significant transition for us, including within our management team.During the first quarter of 2022, Dr. Kadia was appointed our new Chief Executive Officer effective March 9, 2022, and Dr. Hobbs transitioned out of the role of Chief Executive Officer and into the role of President of Global Operations. References to “CEO” in this “Executive Compensation” section shall means Dr. Hobbs before March 9, 2022 and Dr. Kadia on and after March 9, 2022. See “Employment Arrangements” section for additional detail on the terms of the transition.
Mr. Joshi was appointed as our new Chief Financial Officer effective July 25, 2022 and Mr. Chaplin was appointed Chief Legal Officer and Corporate Secretary effective September 19, 2022.

Compensation Philosophy

Our compensation programs are designed to:
Provide a fair, flexible, and market-driven total compensation package,
Attract, motivate, incentivize, and retain the key management team members who contribute to the Company’s short- and long-term success,
Emphasize performance-based pay elements that reward the achievement of business objectives and key results, and
Align the interests of the management team with those of the stockholders by focusing on long-term equity incentives that correlate the growth of sustainable long-term value.

The 2020 amountsCompensation Committee, which is comprised entirely of independent directors, is responsible for assisting our Board of Directors in fulfilling its oversight responsibilities relating to the overall compensation structure, the administration of the Company’s equity, pay policies and programs, and our processes and procedures for the consideration and determination of executive compensation. See the sections titled “Board of Directors Committees” and “Compensation and Leadership Development Committee” beginning on page 8 for more information about the Compensation Committee and how to access its charter.
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Our CEO, CFO, and Chief Human Resources Officer provide our Compensation Committee with information on company and individual performance, competitive practices and perspectives, and preliminary recommendations regarding compensation matters with respect to all executive officers, other than the CEO. No executive officer participates in deliberations regarding their own pay. The Compensation Committee reviews management’s preliminary recommendations and makes final compensation decisions.
The Compensation Committee has the authority to engage its own advisors in carrying out its responsibilities and in connection with the establishment of our executive compensation programs and related policies. In 2022, the Compensation Committee engaged FW Cook as its independent compensation consultant to assist with its duties, including providing objective analysis, input on the compensation peer group selection, insight into competitive compensation practices, and advice and recommendations on executive officer compensation in connection with the Compensation Committee’s decision-making process for 2022. FW Cook did not provide services to us other than the services to the Compensation Committee, as described herein.

Elements of our Executive Compensation Program
The 2022 executive compensation program was designed with our guiding principles in mind, including to attract key talent to lead our company to its new phase of growth and align incentives for our NEOs with the long-term interests of our stockholders. Our 2022 compensation program was comprised of the following three primary elements: 1) base salary, 2) annual bonus and 3) equity awards. Each element is individually and collectively established as part of our program in order to achieve our compensation objectives.

Base Salary
A base salary provides our NEOs with a fixed component of annual compensation for performing specific duties and functions. Base salaries for the NEOs are initially established through arm’s-length negotiation at the time of their hiring, taking into account competitive market compensation paid by other companies for similar positions within the industry, their position, qualifications, experience, prior salary level, and internal parity. Base salaries are reviewed annually, typically in connection with our annual performance review process and may be adjusted from time to time by the Compensation Committee after taking into account independent objective market analyses, individual responsibilities, performance, and experience.
2022 annual base salaries for Dr. Hobbs, Dr. BreinlingerKadia, Mr. Joshi and Mr. Rosinack reflectChaplin were set at the increasetime of hire. No adjustments were made to Dr. Hobbs’ 2022 base salary per his Transition Agreement (which agreement is described in their annual base salaries to $530,450, $387,000 and $275,625 effective February 1, 2020, as well asfurther detail below).
The following table sets forth the further increase in Mr. Rosinack's annual base salary to $295,000 effective September 1, 2020.
(2)The amount shown for Mr. Rosinack represents a discretionary bonus related to activities to support certain internal finance initiatives.
(3)For the option awards column, amounts shown represents the grant date fair value of stock awards and options granted during fiscal years 2019 and 2020 as calculated in accordance with ASC Topic 718. These amounts do not correspond to the actual value that may be recognized by the NEO upon exercise of the applicable awards or sale of the underlying shares of stock. For a discussion of valuation assumptions used to determine the grant date fair value of the stock options granted to our NEOs in fiscal year 2020, see Note 11 of Notes to our Consolidated Financial Statements set forth in our Annual Report on Form 10-K for the year ended December 31, 2020.2022. The actual amount included for 2019 forof base salary paid to our NEOs (other than Dr. Hobbs includesHobbs) was prorated based on length of employment with us in 2022.

Named Executive OfficerBase Salary
Siddhartha Kadia, Ph.D.567,449 
Eric D. Hobbs, Ph.D.560,000 
Mehul Joshi196,901 
Scott Chaplin115,216 

Annual Bonus
An annual bonus provides our NEOs with an optionopportunity to purchase 1,000,000 shares for which vesting is earnedearn additional compensation based on achievement of certainpre-determined short-term corporate goals approved by the Compensation Committee in order to incentivize our executives to drive growth in key areas of our business.
Target annual bonuses for the NEOs are initially established through arm’s-length negotiation at the time of their hiring, taking into the same factors as described above for base salaries. Their target annual bonuses are reviewed annually, typically in connection with our annual performance goals establishedreview process and may be adjusted from time to
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time by the Compensation Committee after taking into account independent objective market analyses, individual responsibilities, performance, and experience.
2022 target annual bonuses for fiscal years 2019Dr. Kadia and Messrs. Joshi and Chaplin were set at the time of hire. No adjustments were made to 2022. The grant date fair value reported assume the probable outcomeDr. Hobbs’ 2022 target annual bonus per his Transition Agreement.
Target annual bonuses are expressed as a percentage of the applicable performance-based vesting conditions.NEO’s base salary received in the year. The grant date fair valuefollowing table sets forth the target annual bonus for our NEOs for the year ended December 31, 2022.

Named Executive OfficerBase Salary ($)Target Annual Bonus as % of Base SalaryTarget Annual Bonus ($)(1)
Siddhartha Kadia, Ph.D.700,000 100 %700,000 
Eric D. Hobbs, Ph.D.560,000 75 %420,000 
Mehul Joshi450,000 50 %225,000 
Scott Chaplin400,000 50 %200,000 
(1) This amount represents the annualized target bonus for 2022. Each NEO’s, except Dr. Kadia’s actual target annual bonus for 2022, was pro-rated based on the length of employment with us in 2022. Dr. Kadia’s 2022 bonus was guaranteed in an amount equal to no less than his pro-rated 2022 target annual bonus, and as such, this was the performance-based shares subjectamount paid to the option assuming the highest level of performance conditions will be achieved is $4,070,065.him for 2022

NarrativeEach of our NEOs participated in our 2022 annual bonus plan, which provided an opportunity for our NEOs and other employees to summary compensation tableearn bonuses based upon our achievement of the following key goals for 2022: GAAP revenue, execution on R&D milestones, year-end cash, and operational goals.

2020 salaries
GoalWeightingThresholdBelow TargetTargetMaximum
0%50%100%125%
GAAP Revenue30%<$80M$85M$100M>$125M
Year-end Cash30%<$115M$115M$125M>$135M

GoalWeightingThresholdMinimumBelow TargetTarget
0%50%75%100%
R&D30%<3 goals3 of 5 goals4 of 5 goals5 of 5 goals
This metric included five goals relating to the delivery of key R&D initiatives.

GoalWeightingThresholdBelow TargetTarget
0%50%100%
Operational10%<2 goals2 of 3 goals3 of 3 goals
This metric included three goals designed to advance our capabilities to establish a data driven culture of decision making.
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Our NEOs each receive a base salary to compensate themIn the first quarter of fiscal year 2023, the Compensation Committee reviewed our achievements for services rendered to our company. The base salary payablethe goals above and determined that we achieved below threshold level for the revenue goal, 93% of the year-end cash goal, 50% of the R&D goal, and 100% of the operational goal.Based on the relative weightings assigned under the 2022 bonus plan, the overall achievement was 53% and bonuses were paid to each named executive officer is intended to provide a fixed component of compensation reflecting the executive’s skill set, experience, role and responsibilities.

The Company increased the base salary of Dr. Hobbs and Mr. Rosinack effective asMessrs. Joshi and Chaplin in an amount equal to 53% of February 1, 2019 to $515,000 and $262,500, respectively. For fiscal year 2020, the Company increasedhis 2022 target bonus amount, pro-rated based on length of employment with us in 2022. Under his employment offer letter, Dr. Hobbs, Dr. Breinlinger and Mr. Rosinack's base salary to $530,450, $387,000 and $275,625 on February 1, 2020, respectively. Mr. Rosinack's base salary was further increased to $295,000 on September 1, 2020 in conjunction with is promotion to Chief Accounting Officer.

Our board of directors and compensation committee may adjust base salaries from time to time in their discretion.

2020 bonuses

For service in fiscal year 2020, there were no formal bonus plans or arrangements applicable to Dr. Hobbs, Dr. Breinlinger and Mr. Rosinack. In the fiscal year 2020, Mr. Rosinack earned a bonus related to individual contributions to certain internal key initiatives that amounted to $24,400. SuchKadia’s 2022 bonus was discretionaryguaranteed in an amount equal to no less than his pro-rated 2022 target annual bonus, and as such, this was the amount paid to him for 2022. The amount of the bonus paid to each NEO under the 2022 bonus plan is set forth in the “Summary Compensation Table” below.
To conserve cash and to further align the interest of the NEOs with our stockholders, the Compensation Committee determined to settle each NEO’s 2022 bonus in the form of fully vested RSUs, with the actual number of RSUs based on senior management determination andthe bonus amount divided by $1.84/share (which was not partthe closing price of any established bonus program.

Equity-based compensation

In February 2020, we granted each of Dr. Hobbs, Dr. Breinlinger and Mr. Rosinack an option to purchase 100,000, 50,000 and 18,000 shares of our common stock, respectively, each with an exercise price pera share equal to the fair market value of our common stock on March 3, 2023, the dategrant date). See footnote 3 in the “Summary Compensation Table” for more information.

Equity Incentives
We view equity-based compensation as a critical component of grant. Eachour balanced total compensation program. Equity-based compensation creates an ownership culture among our employees that provides an incentive to contribute to the continued growth and development of our business and aligns interest of executives with those of our stockholders. In addition, we believe that equity grants promote executive retention because they incentivize our executive officers to remain in our employment during the vesting period.
Historically, we have used stock options and RSU awards as an incentive for long-term compensation to our Named Executive Officers because stock options allow our executive officers to realize value from this form of equity compensation only if our stock price increases, and RSU awards align the interests of our executive officers with the interests of our stockholders generally. Our executives generally are awarded an initial grant in the form of a stock options and/or RSUs, or in the case of Dr. Kadia, performance stock options, in connection with their commencement of employment with us. The Compensation Committee may award additional grants on a periodic basis in order to specifically incentivize executives to achieve certain goals or reward them for exceptional performance.
In 2022, the size and form of equity awards granted to Dr. Kadia and Messrs. Joshi and Chaplin was determined through negotiations at the time of hiring after considering the same factors as described above that were used for setting their base salary. Dr. Kadia’s performance stock options and the terms of that grant were determined with input from FW Cook to provide him with incentives to grow the value of our business over the long-term through stock price performance, further aligning his interest with those of our stockholders.
Dr. Hobbs’s 2022 RSU transition award was determined using an objective and independent market analysis to ensure a smooth transition and motivate Dr. Hobbs in his new role.
The following table is a summary of equity awards granted to our NEOs during 2022:
OptionsRSUsPerformance Options
Name($) (1)(# of shares)(2)($) (1)(# of shares)(2)($) (1)(# of shares)(2)
Siddhartha Kadia, Ph.D.1,157,517339,0595,787,7371,017,1771,714,281.00 678,118 
Eric D. Hobbs, Ph.D.986,265173,333— — 
Mehul Joshi1,250,445451,208 1,249,999287,356 — — 
Scott Chaplin750,081347,910749,998227,963— — 
(1) The dollar values of the options, RSUs and performance options were calculated in accordance with FASB ASC 718. Please see footnote 1 to the “Summary Compensation Table” for additional information.
(2) The equity awards vestsgenerally vest over four years, subject to the NEO’s continued service with us, consistent with our equity grant practices, except for Dr. Kadia’s whose equity awards vest over a special schedule that was negotiated in connection with him joining the Company and becomes exercisableDr. Hobbs whose 2022 RSU transition award
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vesting schedule was negotiated in connection with his transition. Please see the “Outstanding Equity Awards as of December 31, 2022” for additional information as well as the description below with respect to Dr. Kadia’s performance option.

For Dr. Kadia’s performance option, the vesting and exercisability are subject to both a performance-based requirement based upon increases in stock price of the Company and a continued service-based requirement.
The service-based requirement for the performance option is satisfied with respect to 20% of the total number shares subject to the performance option shares on each anniversary of his employment start date, subject to Dr. Kadia’s continued service through the applicable date. This service-based requirement will be fully satisfied in the event Dr. Kadia experiences a “covered termination” (as defined in his Change of Control and Severance Agreement. See “Change of Control and Severance Benefits” section below).
The performance-based requirement will be satisfied as to 1/48th20% of the total number of shares on each monthly anniversary stated vesting start date, subjectsubject to the performance option if the Stock Value is at least equal to the applicable holder’s continued service tostock price goal set forth in the Company throughtable below during the applicable vestingseven-year performance period that commenced on the grant date. Furthermore, we issued Mr. Rosinack an option to purchase 30,000 shares“Stock Value” generally means upon any date during the performance period, the average of our common stock on September 20, 2020 with an exercisethe closing trading price per share equalof the Company’s common stock over the immediately trailing 90-day period. If a “Change in Control” (as defined in the 2020 Plan) occurs before the end of the performance period, Stock Value will mean the per common share dollar amount of cash and the value of any securities or other property paid to the fair market valueholders of ourshares of Company common stock on the date of grant. Such award vests and becomes exercisable as to 1/48th of the shares on each monthly anniversary date of the grant date, subject to Mr. Rosinack's continued service to(or the Company, through the applicable anniversary date.as applicable) as consideration in that Change in Control.

We have adopted a 2020 Incentive Award Plan, referred to below as the 2020 Plan, in order to facilitate the grant of cash
Stock Price GoalNumber of Shares that incrementally vest upon Achievement of a Stock Price Goal
$11.8620% of the total performance option shares
$23.7220% of the total performance option shares
$47.4420% of the total performance option shares
$88.9520% of the total performance option shares
$118.6020% of the total performance option shares
Perquisites and equity incentives to directors, employees (including our named executive officers) and consultants of our company and certain of its affiliates and to enable us to obtain and retain services of these individuals, which is essential to our long-term success. The 2020 Plan was effective on the day prior to our public trading date. For additional information about the 2020 Plan, please see the section titled “Equity compensation plans” below.
Other Elements of CompensationEmployee Benefits
Retirement savings and health and welfare benefits

We currently maintainIn 2022, we provided the following perquisites to Dr. Kadia that were negotiated in connecting with his hiring: (1) $10,000 per month for temporary lodging and travel expenses associated with travel to our headquarters during the first 12 months of employment and (2) legal fees incurred in connection with the negotiations of his new hire employment arrangements, up to a 401(k) retirement savings plan for our employees, including our named executive officers, who satisfy certain eligibility requirements. Our NEOs are eligible to participate in the 401(k) plan on the same terms as other full-time employees. We believe that providing a vehicle for tax-deferred retirement savings though our 401(k) plan adds to the overall desirabilitymaximum of our executive compensation package and further incentivizes our employees, including our named executive officers, in accordance with our compensation policies.$25,000.

All of our full-time employees, including our NEOs, are eligible to participate in our health and welfare plans including medical, dental and vision benefits; medical401(k) plan, which provides for employee elective deferrals and dependent care flexible spending accounts; short-term and long-term disability insurance; and life and AD&D insurance.employer matching contributions.

PerquisitesWe believe the benefits described above are necessary and other personal benefitsappropriate to provide a competitive compensation package to our NEOs and, in the case of Dr. Kadia, to recruit him to join as our CEO.

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TableSpecific Employment and Change of contents
We did not provide any perquisites to our NEOs in fiscal year 2020Control and 2019 that were not provided to all employees generally, but our compensation committee may from time to time approve them in the future when our compensation committee determines that such perquisites are necessary or advisable to fairly compensate or incentivize our employees.
EmploymentSeverance Arrangements with Named Executive Officers
This section contains a description of the material terms of the employment arrangements with our NEOs. Our NEOs signed employment agreements or letters with us, which provide for at-will employment and set forth other terms of employment, including the initial base salary, target incentive opportunity and the terms of the initial equity grants. In addition, each of our NEOs executed a form of our standard confidential information, invention assignment and arbitration agreement.
Offer letters

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Employment Arrangements

Siddhartha Kadia, Ph.D.
In March 2022, we entered into an employment agreement with Dr. Kadia, under which he was entitled to an annual base salary of $700,000, an annual target bonus equal to 100% of his then-current base salary (which bonus was guaranteed and paid on a pro-rata basis at target level for fiscal year 2022), and eligibility to participate in our benefit plans. In addition, Dr. Kadia’s employment agreement provided for certain new hire equity awards as described in greater detail under the “Equity Incentives” section above and perquisites as described in greater detail under the “Perquisites and other Employee Benefits” section above.

Eric D. Hobbs, Ph.D.

In May 2013, we entered into an offer letter with Dr. Hobbs, under which he was entitled to an annual base salary of $200,000, eligibility to participate in our benefit plans and an option to purchase 75,000 shares of our common stock. For
In addition to his employment agreement, we entered into a descriptiontransition agreement with Dr. Hobbs in March 2022 (the “Transition Agreement”). Under this Transition Agreement, Dr. Hobbs agreed to serve as President of Global Operations, or if, so elected by Dr. Hobbs, as a consultant to the Company through December 31, 2023.As an employee of the material terms of this stock option grant, see footnotes 1 and 2Company, pursuant to the outstandingTransition Agreement,
Dr. Hobbs was entitled to compensation at the rate in effect on the date of the agreement, which was an annual base salary of $560,000, an annual target bonus equal to 75% of his annual base salary, and eligibility to participate in our benefit plans. In addition, the agreement provided for a certain transition award as described in greater detail under the “Equity Incentives” section above. Effective for fiscal year 2023, Dr. Hobbs’ annual base salary has been adjusted to $450,000 with an annual target bonus equal to 50% of base salary.
Under the Transition Agreement, if Dr. Hobbs transitions to a consultant, he will be entitled to receive a monthly retainer of $12,000. If the Company terminates Dr. Hobbs’ service arrangement prior to December 31, 2023, Dr. Hobbs will immediately vest in all unvested equity awards at 2020 fiscal year end table.that would have vested as of December 31, 2023.

Keith J. Breinlinger, Ph.D.

Mehul Joshi
In February 2013,July 2022, we entered into an offer letter with Dr. Breinlinger,Mr. Joshi, under which he was entitled to an annual base salary of $200,000,$450,000, an annual target bonus of 50% of his then-current base salary and eligibility to participate in our benefit plans and an option to purchase 90,000 shares of our common stock. For a description of the material terms of this stock option grant, see footnotes 1 and 2 to the outstandingplans. In addition, Mr. Joshi’s offer letter provided for certain new hire equity awards at 2020 fiscal year end table.as described in greater detail under the “Equity Incentives” section above.

Matthew W. Rosinack

Scott Chaplin
In February 2016,September 2022, we entered into an offer letter with Mr. Rosinack,Chaplin, under which he was entitled to an annual base salary of $220,000,$400,000, an annual bonus of 50% of his then-current base salary and eligibility to participate in our benefit plans, $10,000 in the form of a one-time sign-on bonus and an option to purchase 75,000 shares of our common stock. For a description of the material terms of this stock option grant, see footnotes 1 and 2 to the outstandingplans. In addition, Mr. Chaplin’s offer letter provided for certain new hire equity awards at 2020 fiscal year end table.as described in greater detail under the “Equity Incentives” section above.

Change of Control and Severance agreementsBenefits

In July 2020, in connection with our initial public offering, we approved the entry into new severance agreements with all of our named executive officersNEOs that supersedeswere employed as of that date that supersede and replacesreplace the severance benefits they would otherwise be entitled to receive.

Since July 2020, we have entered into severance agreements with each of our NEOs hired after that date.
Under these severance agreements with each of our NEOs, if such NEO’s employment with us is terminated without “cause” or such NEO resigns for “good reason” (as each is defined in the severance agreement), the applicable NEO will be entitled to receive: (i) nine months of continued base salary (or 12 months for Dr. Hobbs)the CEO), (ii) in the case of
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our CEO, an amount equal to his annual bonus for the year of termination assuming 100% of target performance, and (ii)(iii) payment or reimbursement of the cost of continued healthcare coverage for nine months (or 12 months for Dr. Hobbs)the CEO). In lieu of the foregoing benefits, if each NEO’s employment with us is terminated without “cause” or such NEO resigns for “good reason” during the 4545-day period (or 90 day period in the case of our CEO) prior to, or the 12-month period following a Change in Control (as defined in the 2020 Plan), hesuch NEO will be entitled to receive: (i) 12 months of continued base salary (or 18 months for Dr. Hobbs)the CEO), (ii) payment or reimbursement of the cost of continued healthcare coverage for 12 months (or 18 months for Dr. Hobbs)the CEO), (iii) an amount equal to 12 months of such NEO’s annual bonus for the year of termination assuming 100% of target performance (or 18 months for Dr. Hobbs)the CEO) and (iv) full accelerated vesting of any of his unvested equity awards (except for any performance awards). Notwithstanding the foregoing, with respect to Dr. Hobbs’ performance-based stock option described above, all performance conditions subject to the option will be deemed satisfied at target in addition to the accelerated vesting described above, effective as of the date of the qualifying termination.
The foregoing severance benefits are subject to his delivery of an executed release of claims against us and continued compliance with his confidentiality agreement with us.
With respect to Dr. Kadia’s performance option grant, if a change in control of the Company occurs, the percentage of shares that will vest in the table above in the “Equity Incentives” section will be determined using linear interpolation. In addition, even if the stock price goals are not achieved as a result of a Change in Control, at a minimum the vesting of that number of shares subject to the performance option will be accelerated such that the Stock Value Dr. Kadia is eligible to receive with respect to the shares subject to all of the equity awards he received in connection with his hiring in the Change in Control is equal to at least $8,000,000. In the event such Stock Value (after giving effect to full acceleration of the shares subject to the performance option) is less than $8,000,000 in the aggregate, then upon the change in control, the Company will pay to Dr. Kadia a one-time cash bonus equal to the difference between (i) $8,000,000 and (ii) the Stock Value in the change in control with respect to the shares subject to the equity awards granted in connection with his hiring. Notwithstanding the foregoing, in the event Dr. Kadia experiences a covered termination during a “change in control period” (as defined in the severance agreement), then Dr. Kadia shall remain eligible to receive such cash bonus, payable at the closing of the change in control.
Dr. Hobbs is also entitled to certain additional termination benefits under his transition agreement if his service with the Company terminates before December 31, 2023. See “Employment Arrangements” section above.

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Summary Compensation Table
Name and Principal PositionYearSalary
($)
Option
Awards
($)(1)
Stock Awards ($)(2)Non-equity
Incentive Plan
Compensation
($) (3)
All Other Compensation ($)(7)Total
($)
Siddhartha Kadia, Ph.D.(4)
2022567,449 2,871,798 5,787,737 583,333 154,485 9,964,802 
Chief Executive Officer
Eric Hobbs, Ph.D.2022560,020 — 986,265 222,600 13,010 1,781,895 
Former CEO and current Executive Vice President of Integrations and Business Transformation2021557,538 2,281,314 768,196 94,600 15,587 3,717,235 
Mehul Joshi (5)
2022196,901 1,250,445 1,249,999 59,623 9,657 2,766,625 
Chief Financial Officer
Scott Chaplin (6)
2022116,116 750,081 749,998 35,332 774 1,652,301 
Chief Legal Officer
________________

(1)For the options awards column, amounts shown represents the grant date fair value of contentsstock options granted during fiscal years 2021 and 2022 as calculated in accordance with ASC Topic 718. The amount included for Dr. Kadia in 2022 includes an option to purchase 678,118 shares for which vesting is earned based on achievement of certain stock price goals during a 7-year performance period following the grant date. For this grant, the amount was calculated using a model based on multiple stock price paths developed through the use of a Monte Carlo simulation that incorporates into the valuation the possibility that the stock price goals may not be satisfied. These amounts do not correspond to the actual value that may be recognized by the NEO upon exercise of the applicable awards or sale of the underlying shares of stock. For a discussion of valuation assumptions used to determine the grant date fair value of the stock options granted to our NEOs, see Note 13 to our Consolidated Financial Statements set forth in our Annual Report.
(2)The grant date fair value for time-based RSUs is measured in accordance with FASB ASC 718 and based on the closing price of the Company’s common stock on the date of grant.
(3)Represents amounts paid under the Company's annual bonus plan as further described under “Executive Compensation.” Annual incentives are based on the performance of predetermined goals for the year. Annual bonuses for fiscal year 2022 were not paid in cash and instead the Company issued fully vested RSUs on March 3, 2023 with a grant date fair value of $1.84 per share. The number of RSUs issued to our NEOs in settlement of their 2022 annual bonuses was: 317,029 (Dr. Kadia), 120,978 (Dr. Hobbs), 32,404 (Mr. Joshi), and 19,902 (Mr. Chaplin).
(4)Dr. Kadia was appointed on March 9, 2022 as our Chief Executive Officer.
(5)Mr. Joshi was appointed on July 24, 2022 as our Chief Financial Officer.
(6)Mr. Chaplin was appointed on September 19, 2022 as our Chief Legal Officer.
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(7)Amounts represent (i) the Company’s 401(k) matching contributions to Dr. Kadia’s account in the amount of $12,200 for fiscal year 2022, director fees earned as a non-employee director in the amount of $16,250 for fiscal year 2022, housing allowance of $100,000 for fiscal year 2022, reimbursement of $25,000 of legal fees in connection with Dr. Kadia’s employment agreement and term life insurance premiums paid by the Company in the amount of $1,035 for fiscal year 2022; (ii) the Company’s 401(k) matching contributions to Dr. Hobbs account in the amount of $12,200 for fiscal year 2022 and $14,987 for fiscal year 2021 and term life insurance premiums paid by the Company in the amount of $810 for fiscal year 2022 and $600 for fiscal year 2021; (iii) the Company’s 401(k) matching contributions to Mr. Joshi’s account in the amount of $7,875 for fiscal year 2022 and term life insurance premiums paid by the Company in the amount of $1,782 for fiscal year 2022; (iv) term life insurance premiums paid for Mr. Chaplin in the amount of $774 for fiscal year 2022.




































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Outstanding Equity Awards as of December 31, 20202022
The following table lists the outstanding equity awards held by our NEOs as of December 31, 2020:2022:
Option awards
NameVesting commencement date (1)(2)Number of
securities
underlying
unexercised
options
(#) exercisable
Numbers of
securities
underlying
unexercised
 options
(#) unexercisable
Equity
incentive
plan
awards:
number of
securities
underlying
unexercised
unearned
options
(#)

Option
exercise
price
($)
 Option
expiration
date
Eric D. Hobbs, Ph.D.8/16/20161,3020.64 10/13/2024
Chief Executive Officer1/6/2015(3)(4)37,5000.64 2/12/2025
3/24/2016(4)60,0002.26 3/23/2026
4/14/2017229,16720,8322.54 4/13/2027
11/2/2017385,422114,5772.10 11/1/2027
2/7/2019(4)458,340541,6606.26 2/6/2029
9/12/2019(5)750,0008.90 9/11/2029
2/6/2020(4)20,83679,16411.06 3/11/2030
Keith J. Breinlinger, Ph.D.3/18/201345,0000.12 6/16/2023
Chief Technology2/14/2015(3)(4)50,0000.12 2/26/2024
Officer8/16/2016(4)50,0000.64 10/13/2024
1/16/201525,0000.64 2/12/2025
3/24/201637,5002.26 3/23/2026
2/2/2017(4)119,7925,2082.54 2/1/2027
11/2/2017(4)115,62634,3742.10 11/1/2027
2/7/2019(4)114,588135,4116.26 2/6/2029
2/1/2020(4)10,42039,58011.06 3/3/2030
Matthew W. Rosinack3/21/201641,0412.50 5/31/2026
Chief Accounting3/1/2017(4)9,3766242.54 4/4/2027
Officer1/18/2018(4)11,09710,1532.24 1/17/2028
2/7/2019(4)8,1189,5906.26 2/6/2029
2/1/2020(4)3,75014,25011.06 3/10/2030
9/20/2020(4)1,87528,12572.31 9/19/2030
________________
Option awardsStock Awards
NameVesting commencement date (1)Number of
securities
underlying
unexercised
options
(#) exercisable
Numbers of
securities
underlying
unexercised
 options
(#) unexercisable(a)
Equity
incentive
plan
awards:
number of
securities
underlying
unexercised
unearned
options
(#)

Option
exercise
price
($)
 Option
expiration
date
Number of Shares or Units of Stock That Have Not Vested (#)Market value of Shares or Units of Stock that Have Not Vested ($)(b)
Siddhartha Kadia, Ph.D.3/10/2022(2)84,765254,2945.69 3/9/2032— — 
3/10/2022(3)678,1185.69 3/9/2032— — 
9/7/2021(4)2,0412,85937.35 9/6/2031— — 
9/7/2021(5)— — 2,858 7,659 
3/10/2022(2)— — 762,883 2,044,526 
Eric Hobbs, Ph.D.11/2/2017(6)182,4992.10 11/1/2027— — 
3/24/2016(6)60,0002.26 3/23/2026— — 
4/14/2017(7)126,0322.54 4/13/2027— — 
2/7/2019(7)958,33341,6676.26 2/6/2029— — 
12/31/2021(9)125,0008.90 9/12/2029— — 
12/31/2020(9)83,3338.90 9/12/2029— — 
2/6/2020(7)70,83729,16311.06 3/11/2030— — 
3/4/2021(7)37,45048,15053.72 3/1/2031— — 
3/4/2021(8)— — 8,044 21,558 
3/10/2022(2)— — 43,333 116,132 
Mehul Joshi7/25/2022(6)451,0284.35 7/25/2032— — 
7/25/2022(10)— 0287,356 770,114 
Scott Chaplin9/19/2022(6)347,9103.29 9/18/2032— — 
9/19/2022(11)— — 227,963 610,941 
(1)ExceptThe dollar amounts shown in column (b) is determined by multiplying the number of shares or units shown in column (a), as otherwise noted,applicable, by $2.68, the closing price of a share of the Company’s common stock on December 30, 2022, the last trading day of the Company’s fiscal year. The stock options and RSUs held by our NEOs are eligible to receive accelerated vesting, as described above under their severance agreements. See “Executive Compensation—Change of Control and Severance Benefits” section above for a more detailed description of the stock awardsprice goals.
(2)1/12th of the shares vest ason each quarterly anniversary of the grant date subject to the holder continuing to provide services to the Company through such vesting date.
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(3)The service-based requirement for the performance options will be satisfied with respect to 20% of the total number of shares underlying the performance options on each anniversary of the appointment date subject to Dr. Kadia continuing to provide services to the Company through such vesting date. The performance-based requirement will be satisfied in respect of that percentage of the total number of shares subject to the performance options determined based on certain stock price goals, whereby any such stock price goal must be achieved by the seventh anniversary of the appointment date. See “Executive Compensation – Equity Incentives” for a more detailed description of the stock price goals.
(4)1/36th of the shares vest monthly subject to the holder continuing to provide services to the Company through such vesting date.
(5)33% of the shares vest on the one-year anniversary of the vesting commencement date and vest quarterly thereafter for two years, subject to the holder continuing to provide services to the Company through such vesting date.
(6)25% of the shares vest on the one yearone-year anniversary of the vesting commencement date and vest as to 1/48th of the shares monthly thereafter, such that all awards will be vested on the four yearfour-year anniversary of the vesting commencement date, subject to the holder continuing to provide services to the Company through such vesting date.
(2)All of the stock options held by each of our NEOs are eligible to receive accelerated vesting, as described above under their severance agreements.
(3)The option is early exercisable in exchange for restricted shares subject to a right of repurchase in favor of the Company.
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(4)(7)1/48th of the shares vest on each monthly anniversary of the vesting commencement date, such that all awards will be vested on the four yearfour-year anniversary of the vesting commencement date, subject to the holder continuing to provide services to the Company through such vesting date.
(5)(8)1/16th of the shares vest on each quarterly anniversary of the vesting commencement date, such that all awards will be vested on the four-year anniversary of the vesting commencement date, subject to the holder continuing to provide services to the Company through such vesting date.
(9)The shares become eligible to vest based on the achievement of certain performance goals established for fiscal years 2019 to 2022, which were established and evaluated by our board of directors. Vesting of the applicable number of shares subject to the option will occuroccurred in substantially equal installments over a 12-month or 24-month period (depending upon the specific performance criteria) upon the achievement of the applicable performance goal, subject to Dr. Hobbs’ continued service through the applicable vesting date, and the shares subject to the option are either achieved or not achieved.date. As of December 31, 2020, one2022, two performance criteria had been met and 750,000 suchthe other seven performance criteria were not achieved and thus forfeited. The options remained outstanding.associated with the two achieved performance criteria were fully vested as of December 31, 2022.
(10)25% of the shares vest on the one-year anniversary and vest as to 1/16th of the shares monthly thereafter, such that all awards will be vested on the four-year anniversary of the vesting commencement date, subject to the holder continuing to provide services to the Company through such vesting date.
(11)25% of the shares vest on August 20, 2023 with the remaining awards vesting in equal installments over the next nine quarters, subject to the holder continuing to provide services to the Company through such vesting date.














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Pay Versus Performance
As required by Section 953(a) of the Dodd-Frank Wall Street Reform and Consumer Protection Act, and Item 402(v) of Regulation S-K, we are providing the following information about the relationship between executive compensation actually paid and certain financial performance of the Company. As a smaller reporting company, we are only required to provide the following information for the two most recent fiscal years.
Year (a)Summary Compensation Table Total for CEO- Hobbs (b) (1)Compensation Actually Paid to CEO- Hobbs (c)(4)Summary Compensation Table Total for CEO-Kadia (d) (2)Compensation Actually Paid to CEO- Kadia (e) (4)Average Summary Compensation Table Total for Non-CEO NEOs (f)(3)Average Compensation Actually Paid to Non-CEO NEOs (g) (4)Value of Initial Fixed $100 Investment Based on Total Shareholder Return (h) (5)Net Loss (i) (6)
2022$1,781,895 $(6,102,013)$9,964,802 $5,345,487 $2,210,463 $1,463,884 $3.00 $(98,040,000)
2021$3,717,235 $(51,477,577)N/AN/A$1,765,473 $(5,441,496)$20.33 $(71,724,000)
(1) Represents the amount of total compensation reported for Dr. Hobbs (our former Chief Executive Officer and current Executive Vice President of Integrations and Business Transformation for each corresponding fiscal year in the “Total” column of the Summary Compensation Table. Refer to “Executive Compensation – Summary Compensation Table.”
(2) Represents the amount of total compensation reported for Dr. Kadia (our current Chief Executive Officer) for 2022 in the “Total” column of the Summary Compensation Table. Refer to “Executive Compensation – Summary Compensation Table.”
(3) Represents the average of the amounts reported for the Company’s named executive officers as a group (excluding Drs. Kadia and Hobbs for 2022 and Dr. Hobbs for 2021) in the “Total” column of the Summary Compensation Table in each applicable year. Refer to “Executive Compensation – Summary Compensation Table.” The named executive officers (excluding Drs. Kadia and Hobbs for 2022 and Dr. Hobbs for 2021) included for purposes of calculating the average amounts in each applicable year are: (i) for 2022, Mehul Joshi (our Chief Financial Officer) and Scott Chaplin (our Chief Legal Officer); and (ii) for 2021, Kurt Wood (our former Chief Financial Officer), Shaun M. Holt (our former Chief Financial Officer), Keith Breinlinger, Ph.D. (our former Chief Technology Officer), Stuart Merkadeau (our former General Counsel) and Matthew Rosinack (our former Chief Accounting Officer).
(4) Represents the amount of “compensation actually paid” to the respective NEO(s), as computed in accordance with Item 402(v) of Regulation S-K. The amounts do not reflect the actual amount of compensation earned or paid to the respective NEO(s). Adjustments to calculate “compensation actually paid” (columns (c), (e), and (g)) are described and calculated below. For each amount in the “Other NEOs” columns, the amounts relate to an average of the amounts applicable to the respective NEOs discussed in footnote (3) above.
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20222021
AdjustmentsCEO- HobbsCEO- KadiaOther NEOsCEO-HobbsOther NEOs
Summary Compensation Table (“SCT”)Total$1,781,895 $9,964,802 $2,210,463 $3,717,235 $1,765,473 
Adjustments for stock awards and option awards
(Deduct): Aggregate value for stock awards and option awards included in SCT Total for the covered fiscal year(986,265)(8,659,535)(2,000,262)(3,049,510)(1,379,772)
Add: Fair value at year end of awards granted during the covered fiscal year that were outstanding and unvested at the covered fiscal year end116,132 2,976,216 1,253,683 552,094 288,183 
Add (Deduct): Year-over Year change in fair value at covered fiscal year end of awards granted in any prior fiscal year that were outstanding and unvested at the covered fiscal year end(1,209,349)(63,030)— (23,968,674)(2,337,505)
Add: Vesting date fair value of awards granted and vesting during the covered fiscal year520,867 1,165,982 — 317,307 82,538 
Add (Deduct): Change as of the vesting date (from the end of the prior fiscal year) in fair value of awards granted in any prior fiscal year for which vesting conditions were satisfied during the covered fiscal year(4,802,414)(38,949)— (18,935,078)(1,769,639)
(Deduct): Fair value at end of prior fiscal year of awards granted in any prior fiscal year end that failed to meet the applicable vesting conditions during the covered fiscal year(1,522,879)— — (10,110,950)(2,090,773)
Compensation Actually Paid Amounts (as calculated)$(6,102,013)$5,345,486 $1,463,884 $(51,477,576)$(5,441,495)

(5) Represents cumulative Company total shareholder return (TSR). TSR is calculated by dividing the sum of the cumulative amount of dividends, if any, for each measurement period (2021 and 2021-2022), assuming dividend reinvestment, and the difference between the Company’s share price at the end and the beginning of the measurement period by the Company’s share price at the beginning of the measurement period. The Company did not issue any dividends during 2021 or 2022.
(6) Represents the amount of net income reflected in the Company’s audited financial statements for the applicable year.







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chart-1c6cfff53d654d34bd6a.jpg



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Limitation of Liability and Indemnification
Our amended and restated certificate of incorporation and our amended and restated bylaws provide that we will indemnify our directors and officers to the fullest extent permitted under Delaware law, which prohibits our amended and restated certificate of incorporation from limiting the liability of our directors for the following:
any breach of the director’s duty of loyalty to us or our stockholders;
acts or omissions not in good faith or that involve intentional misconduct or a knowing violation of law;
unlawful payments of dividends or unlawful stock repurchases or redemptions; or
any transaction from which the director derived an improper personal benefit.

Our amended and restated certificate of incorporation and our amended and restated bylaws also provide that if Delaware law is amended to authorize corporate action further eliminating or limiting the personal liability of a director, then the liability of our directors will be eliminated or limited to the fullest extent permitted by Delaware law, as so amended. This limitation of liability does not apply to liabilities arising under the federal securities laws and does not affect the availability of equitable remedies such as injunctive relief or rescission.

Our amended and restated certificate of incorporation and our amended and restated bylaws also provide that we shall have the power to indemnify our employees and agents to the fullest extent permitted by law. Our amended and restated bylaws also permit us to secure insurance on behalf of any officer, director, employee or other agent for any liability arising out of his or her actions in this capacity, regardless of whether we would have the power to indemnify such person against such expense, liability or loss under the General Corporation Law of the State of Delaware. We have obtained directors’ and officers’ liability insurance.

We have entered into separate indemnification agreements with our directors and executive officers, in addition to indemnification provided for in our amended and restated certificate of incorporation and amended and restated bylaws.

These agreements, among other things, provide for indemnification of our directors and executive officers for expenses, judgments, fines and settlement amounts incurred by this person in any action or proceeding arising out of this person’s services as a director or executive officer or at our request. We believe that these provisions in our amended and restated certificate of incorporation and amended and restated bylaws and indemnification agreements are necessary to attract and retain qualified persons as directors and executive officers.

The above description of the indemnification provisions of our amended and restated certificate of incorporation, our amended and restated bylaws and our indemnification agreements is not complete and is qualified in its entirety by reference to these documents, each of which is filed as an exhibit to this registration statement to which this prospectus forms a part. The limitation of liability and indemnification provisions in our amended and restated certificate of incorporation and amended and restated bylaws may discourage stockholders from bringing a lawsuit against directors for breach of their fiduciary duties. They may also reduce the likelihood of derivative litigation against directors and officers, even though an action, if successful, might benefit us and our stockholders. A stockholder’s investment may be harmed to the extent we pay the costs of settlement and damage awards against
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directors and officers pursuant to these indemnification provisions. Insofar as indemnification for liabilities under the Securities Act may be permitted to directors, officers or persons controlling us pursuant to the foregoing provisions, we have been informed that in the opinion of the SEC such indemnification is against public policy as expressed in the Securities Act and is therefore unenforceable.

There is no pending litigation or proceeding naming any of our directors or officers as to which indemnification is being sought, nor are we aware of any pending or threatened litigation that may result in claims for indemnification by any director or officer.
Equity Compensation Plan Information
The following table provides information as of December 31, 2020 with respect to shares of our common stock that may be issued under our existing equity compensation plans.
Plan CategoryNumber of Securities
to be Issued upon
Exercise of
Outstanding
Options, Restricted
Stock Units and
Rights
Weighted Average
Exercise Price of
Outstanding Options
and Rights
Number of Securities
Remaining Available
for Future Issuance
Under Equity
Compensation Plans
(Excluding Securities
Reflected in the first
Column)
Equity compensation plans approved by security holders
2020 Incentive Award Plan (1)
1,119,000 $43.41 5,797,516 
2011 Equity Incentive Plan (2)
8,620,303 $5.59 — 
2020 Employee Stock Purchase Plan (3)
— — 612,150 
Equity compensation plans not approved by security holders— — — 
TOTAL9,739,303 $9.94 6,409,666 
________________
(1)Our board of directors adopted, and our stockholders approved, the 2020 Incentive Award Plan (the “2020 Plan”). The initial number of shares authorized and available for issuance in the connection with grant of future awards was 6,750,000. The number of shares remaining for issuance under the 2020 Plan includes awards granted and outstanding under the 2011 Plan that are forfeited or lapse unexercised after the effective date of the 2020 Plan. In addition, the 2020 Plan contains an “evergreen” provision, pursuant to which the number of shares of common stock reserved for issuance pursuant to awards under such plan shall be increased on the first day of each year beginning in 2021 and ending in 2030 equal to the lesser of (A) four percent (4%) of the shares of stock outstanding (on an as converted basis) on the last day of the immediately preceding fiscal year and (B) such smaller number of shares of stock as determined by our board of directors; provided, however, that no more than 50,000,000 shares of stock may be issued upon the exercise of incentive stock options.
(2)Following the adoption of the 2020 Incentive Award Plan in July 2020, any awards outstanding under the 2011 Equity Incentive Plan continue to be governed by their existing terms but no further awards may be granted under the 2011 Incentive Award Plan.
(3)The maximum number of shares of our common stock which are authorized for sale under the ESPP is equal to the sum of (a) 612,150 shares of common stock and (b) an annual increase on the first day of each year beginning in 2021 and ending in 2030, equal to the lesser of (i) 1% of the shares of common stock outstanding (on an as converted basis) on the last day of the immediately preceding fiscal year and (ii) such number of shares of common stock as determined by our board of directors; provided, however, no more than 10,000,000 shares of our common stock may be issued under the ESPP. The shares reserved for issuance under the ESPP may be authorized but unissued shares or reacquired shares. As of December 31, 2020 no shares had been issued under the ESPP.
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STOCK OWNERSHIP

The following table sets forth certain information with respect to the beneficial ownership of our common stock as of March 23, 202131, 2023 for:
each person, or group of affiliated persons, known by us to beneficially own more than 5% of our common stock; 
each named executive officer;
each of our current directors; and
all of our current directors and executive officers as a group.
Beneficial ownership is determined in accordance with the rules of the SEC and generally includes voting or investment power with respect to securities. Unless otherwise indicated below, to our knowledge, the persons and entities named in the table have sole voting and sole investment power with respect to all shares beneficially owned, subject to community property laws where applicable. Common stock subject to options or other rights to acquire common stock that are currently exercisable or exercisable within 60 days of March 23, 202131, 2023 are deemed to be outstanding and to be beneficially owned by the person holding such options or rights for the purpose of computing the percentage ownership of that person, but are not treated as outstanding for the purpose of computing the percentage ownership of any other person.
The percentage of shares beneficially owned is computed on the basis of 66,278,21398,744,915 shares of our common stock outstanding as of March 23, 2021.31, 2023. Shares of our common stock that a person has the right to acquire within 60 days of March 23, 202131, 2023 are deemed outstanding for purposes of computing the percentage ownership of the person holding such rights, but are not deemed outstanding for purposes of computing the percentage ownership of any other person, except with respect to the percentage ownership of all directors and executive officers as a group. Unless otherwise noted below, no selling stockholder has had any position, office or other material relationship with us or any of our predecessors or affiliates within the past three years. Unless otherwise indicated in the footnotes below, based on the information provided to us by or on behalf of the selling stockholders, no selling stockholder is a broker-dealer or an affiliate of a broker-dealer.
Holders of our common stock are entitled to one vote for each share held on all matters submitted to a vote of stockholders.
Unless otherwise indicated, the address for each listed stockholder is: c/o Berkeley Lights,PhenomeX Inc., 5858 Horton Street, Suite 320, Emeryville, California 94608. To our knowledge, except as indicated in the footnotes to this table and pursuant to applicable community property laws, the persons named in the table have sole voting and investment power with respect to all shares of common stock.
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Shares Beneficially Owned
Name and address of beneficial ownerNumberPercent
Directors and Named Executive Officers:
Eric D. Hobbs, Ph.D. (1)
1,300,6431.9 %
Keith J. Breinlinger, Ph.D. (2)
656,1491.0 %
Matthew W. Rosinack (3)
83,779*
Sarah Boyce (4)
90,000*
Igor Khandros, Ph.D. (5)
9,711,62814.7 %
Michael Marks (6)
1,302,4082.0 %
Michael Moritz— *
Elizabeth Nelson (7)
157,500*
James Rothman, Ph.D. (8)
195,720*
Gregory T. Lucier (9)
48,768*
All executive officers and directors as a group (12 persons)13,938,83020.3 %
Shares Beneficially Owned
Name and address of beneficial ownerNumberPercent
Directors and Named Executive Officers:
Scott Chaplin (1)
19,902*
John Chiminski (2)
76,356*
Eric D. Hobbs, Ph.D. (3)
1,948,4041.9 %
Jessica Hopfield, Ph.D. (4)
143,647*
Mehul Joshi (5)
19,586*
Siddhartha Kadia, Ph.D. (6)
675,863*
Igor Khandros, Ph.D. (7)
9,043,0219.2 %
Gregory T. Lucier (8)
302,965*
Michael Moritz (9)
211,820 *
Elizabeth Nelson (10)
215,623*
James Rothman, Ph.D. (11)
358,468*
Peter Silvester (12)
64,596 *
All executive officers and directors as a group (13 persons)13,124,06513.0 %
Shares Beneficially OwnedShares Beneficially Owned
Name and address of beneficial ownerName and address of beneficial ownerNumberPercentName and address of beneficial ownerNumberPercent
5% Stockholders:
5% Stockholders:5% Stockholders:
Entities affiliated with Igor Khandros, Ph.D. (13)
Entities affiliated with Igor Khandros, Ph.D. (13)
9,043,021 9.2 %
ARK Investment Management LLC (14)
ARK Investment Management LLC (14)
8,006,435 8.1 %
Northpond Ventures LLC (15)
Northpond Ventures LLC (15)
5,390,820 5.5 %
Entities affiliated with Igor Khandros, Ph.D. (10)
9,711,628 14.7 %
Entities affiliated with Sequoia Capital (11)
4,828,808 7.3 %
________________
*Less than 1%.
(1)Consists of (i) 186,19819,902 shares of common stock directly held by Dr. Hobbs andMr. Chaplin.
(2)Consists of (i) 20,858 shares of common stock directly held by Mr. Chiminski, (ii) 1,114,45535,524 shares of common stock that may be acquired pursuant to the exercise of stock options within 60 days of March 23, 2021.31, 2023 and (iii) 19,974 RSUs that will vest within 60 days of March 31, 2023.

(2)(3)Consists of (i) 34,536243,029 shares of common stock directly held by Dr. Breinlinger andHobbs (ii) 621,6131,704,482 shares of common stock that may be acquired pursuant to the exercise of stock options within 60 days of March 23, 2021.31, 2023 and (iii) 893 RSUs held that will best within 60 days of March 31, 2023. Dr. Hobbs resigned from the Board on March 9, 2022.

(3)(4)Consists of (i) 17,58684,030 shares of common stock directly held by Mr. Rosinack andDr. Hopfield, (ii) 66,19340,052 shares of common stock that may be acquired pursuant to the exercise of stock options within 60 days of March 23, 2021.31, 2023 and (iii) 19,565 RSUs that will vest within 60 days of March 31, 2023.

(4)(5)Consists of 90,00019,586 shares of common stock directly held by Mr. Joshi.
(6)Consists of (i) 560,121 shares of common stock directly held by Dr. Kadia, (ii) 115,742 shares of common stock that may be acquired pursuant to the exercise of stock options within 60 days of March 23, 2021.31, 2023.

(5)(7)Consists of 9,711,628(i) 8,986,298 shares of common stock directly held by family trusts of which Dr. Khandros is a trustee.

(6)Consists of 1,302,408 shares of common stock directly held by Michael Marks and through various accounts over which Michael Marks exercises investment control.

(7)Consists of (i) 50,000 shares of common stock directly held by Ms. Nelson andtrustee, (ii) 107,50037,158 shares of common stock that may be acquired pursuant to the exercise of stock options within 60 days of March 23, 2021.31, 2023 and (iii) 19,565 RSUs that will vest within 60 days of March 31, 2023.

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(8)Consists of (i) 138,845155,097 shares of common stock directly held by Dr. Rothman andMr. Lucier, (ii) 56,875128,303 shares of common stock that may be acquired pursuant to the exercise of stock options within 60 days of March 23, 2021.

31, 2023 and (iii) 19,565 RSUs that will vest within 60 days of March 31, 2023.
(9)Consists of (i) 22,727155,097 shares of common stock directly held directly by an estate planning vehicle over which Mr. Lucier andMoritz exercises investment control, (ii) 26,04137,158 shares of common stock that may be acquired pursuant to the exercise of stock options within 60 days of March 23, 2021.

31, 2023 and (iii) 19,565 RSUs that will vest within 60 days of March 31, 2023.
(10)Consists of 9,711,628(i) 56,400 shares of common stock directly held by Ms. Nelson, (ii) 139,658 shares of common stock that may be acquired pursuant to the exercise of stock options within 60 days of March 31, 2023 and (iii) 19,565 RSUs that will vest within 60 days of March 31, 2023.
(11)Consists of (i) 144,245 shares of common stock directly held by Dr. Rothman, (ii) 194,658 shares of common stock that may be acquired pursuant to the exercise of stock options within 60 days of March 31, 2023 and (iii) 19,565 RSUs that will vest within 60 days of March 31, 2023.
(12)Consists of 64,596 shares of common stock directly held by Mr. Silvester.
(13)Consists of (i) 8,986,298 shares of common stock directly held by family trusts of which Dr. Khandros is a trustee.trustee, (ii) 37,158 shares of common stock that may be acquired pursuant to the exercise of stock options within 60 days of March 31, 2023 and (iii) 19,565 RSUs that will vest within 60 days of March 31, 2023

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(11)(14)Consists of (i) 1,448,6408,006,435 shares of common stock directly held by SC U.S Venture XVARK Investment Management L.P,LLC (“ARK”), over which ARK has sole voting and dispositive power, as reported by ARK on a Schedule 13G/A filed with the SEC on February 9, 2022. The principal business address of which 1,196,143ARK is 3 East 28th Street, 7th Floor, New York, NY 10016.
(15) Consists of (i) 4,117,272 shares of common stock are directly ownedheld by Sequoia Capital U.S. Venture Fund XV, L.P.Northpond Ventures, LP (“Northpond Ventures”), 18,107 shares are directly owned by Sequoia Capital U.S. Partners Fund XV, L.P., 50,413and (ii) 1,273,548 shares of common stock are directly ownedheld by SequoiaNorthpond Capital, U.S. Venture Partners Fund XV (Q)LP (“Northpond Capital”), L.P.as reported on a Schedule 13G jointly filed by Northpond Ventures, Northpond Capital, Northpond Ventures GP, LLC, Northpond Capital GP, LLC and 183,977Michael P. Rubin filed with the SEC on March 29, 2023. Northpond Ventures GP, LLC (“Northpond Ventures GP”) is the general partner of Northpond Ventures. Northpond Capital GP, LLC (“Northpond Capital GP”) is the general partner of Northpond Capital. Mr. Rubin is the sole managing member Northpond Ventures GP and Northpond Capital GP. As a result of the foregoing relationships, each of Northpond Ventures GP and Mr. Rubin have shared dispositive and voting power over and may also be deemed to beneficially own the shares of common stock are directly ownedheld by SequoiaNorthpond Ventures, and each of Northpond Capital U.S. Venture Principals Fund, L.P.GP and (ii) 3,380,168Mr. Rubin have shared dispositive and voting power over and may also be deemed to beneficially own the shares of common stock are directly held by SC U.S Growth VI Management, L.P.,Northpond Capital. The address of which 3,327,874 shares of common stock are directly owned by Sequoia Capital U.S. Growth Fund VI, L.P. and 142,294 shares of common stock are directly owned by Sequoia Capital U.S. Growth Principals VI Fund, L.P. The general partner of Sequoia Capital U.S. Venture Fund XV, L.P., Sequoia Capital U.S. Partners Fund XV, L.P., Sequoia Capital U.S. Venture Partners Fund XV (Q), L.P. and Sequoia Capital U.S. Venture XV Principals Fund, L.P.the entities mentioned in this footnote is SC U.S. Venture XV Management, L.P. The general partner of Sequoia Capital U.S. Growth Fund VI, L.P. and Sequoia Capital U.S. Growth Principals Fund, L.P. is SC.U.S. Growth VI Management, L.P.7500 Old Georgetown Road, Suite 850, Bethesda, MD 20814.




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Equity Compensation Plan Information
The following table provides information as of contentsDecember 31, 2022 with respect to shares of our common stock that may be issued under our existing equity compensation plans.
Plan CategoryNumber of Securities
to be Issued upon
Exercise of
Outstanding
Options, Restricted
Stock Units and
Rights
Weighted Average
Exercise Price of
Outstanding Options
and Rights
Number of Securities
Remaining Available
for Future Issuance
Under Equity
Compensation Plans
(Excluding Securities
Reflected in the first
Column)
Equity compensation plans approved by security holders
2020 Incentive Award Plan (1)
8,031,812 $10.12 12,690,281 
2011 Equity Incentive Plan (2)
4,183,930 $6.21 — 
2020 Employee Stock Purchase Plan (3)
— — 1,619,235 
Equity compensation plans not approved by security holders— — — 
TOTAL12,215,742 $8.03 14,309,516 
________________
(1)Our board of directors adopted, and our stockholders approved, the 2020 Incentive Award Plan (“2020 Plan”) in July 2020. The initial number of shares authorized and available for issuance under the 2020 Plan was 6,750,000. The number of shares remaining for issuance under the 2020 Plan includes awards granted and outstanding under the 2011 Equity Incentive Plan (“2011 Plan”) that were forfeited or lapsed after the effective date of the 2020 Plan. In addition, the 2020 Plan contains an “evergreen” provision, pursuant to which the number of shares of common stock reserved for issuance pursuant to awards granted under the 2020 Plan shall be increased on the first day of each year beginning in 2021 and ending in 2030 equal to the lesser of (A) four percent (4%) of the shares of stock outstanding (on a converted basis) on the last day of the immediately preceding fiscal year and (B) such smaller number of shares of stock as determined by our board of directors; provided, however, that no more than 50,000,000 shares of stock may be issued upon the exercise of incentive stock options.
(2)Following the adoption of the 2020 Plan, any awards outstanding under the 2011 Plan continue to be governed by their existing terms but no further awards may be granted under the 2011 Plan. Forfeited stock options issued under the 2011 Plan will automatically be added to the 2020 Plan.
(3)Our Employee Stock Purchase Plan (“ESPP”) provides that the number of shares under the ESPP will be increased annually on the first day of each year beginning in 2021 and ending in 2030, equal to the lesser of (i) one percent (1%) of the shares of common stock outstanding (on an as converted basis) on the last day of the immediately preceding fiscal year and (ii) such number of shares of common stock as determined by our board of directors; provided, however, that no more than 10,000,000 shares of our common stock may be issued under the ESPP. The shares reserved for issuance under the ESPP may be authorized but not unissued shares or reacquired shares.
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CERTAIN RELATIONSHIPS AND RELATED PARTY TRANSACTIONS

The following is a description of each transaction since January 1, 20202022 and each currently proposed transaction in which we have been, or are to be, a participant, in which the amount involved exceeded or will exceed $120,000, and in which any of our directors, executive officers or holders of more than 5% of our capital stock, or an affiliate or immediate family member thereof, had or will have a direct or indirect material interest.
Advisory Agreement with James Rothman, Ph.D.
In April 2017, we entered into a strategic/scientific advisor consulting agreement with James E. Rothman, Ph.D., a member of our board of directors, pursuant to which Dr. Rothman agreed to provide us with consulting and advisory services relating to, among other things, our research and development activities and commercialization of our platform. In consideration of Dr. Rothman’s services, we agreed to pay Dr. Rothman $250,000 per year, reimburse his out of pocketout-of-pocket expenses and grant him an option to purchase 250,000 shares of our common stock. The original term of the agreement was for a period of 36 months. In March 2020, we entered into an amendment to the agreement, pursuant to which we agreed to extend the term of the agreement for an additional 36 months (expiring on April 1, 2023), to pay Dr. Rothman $125,000 per year during the extended term, and to grant Dr. Rothman an option to purchase an additional 157,500 shares of our common stock.
Distribution Agreement with Nikon Corporation
In January 2018, we entered into a distribution agreement with Nikon Instech Co., Ltd. and Nikon Corporation, or Nikon, as our exclusive distributor of our products in Japan. Nikon is the holder of greater than 5% of our capital stock, and Makoto Shintani, who was a member of our board of directors from May 2018 until July 2020, was then a corporate vice president at Nikon. In March 2019, we entered into an amended and restated distribution agreement with Nikon, as our exclusive distributor of our products in Japan, Singapore, Thailand and South Korea and our non-exclusive distributor in China. As the exclusive distributor in Japan, Singapore, Thailand and South Korea, Nikon is required to purchase a minimum quantity of our products every six months throughout the term of the distribution agreement, which expires in March 2022. The revenue recognized by us under this distribution agreement during the year ended 2020 was $4,015,835. The accounts receivable under the distribution agreement at December 31, 2020 was $164,379.
Participation in Our Initial Public Offering
Gregory Lucier, a member of our board of directors, purchased 22,727 shares of our common stock for a purchase price of approximately $500,000 in our initial public offering at the public offering price and on the same terms as the other purchasers in our initial public offering.
Amended and Restated Investors’ Rights Agreement
We are party to an amended and restated investors’ rights agreement or the (“Investors’ Rights Agreement,Agreement”), with certain holders of 5% or more of our capital stock and entities affiliated with certain of our directors, as well as certain of our directors and executive officers. The Investors’ Rights Agreement grants rights to certain holders, including certain registration rights with respect to the registrable securities held by them.
Stock Option Grants to Directors and Executive Officers
We have granted stock options to our directors and executive officers. For more information regarding the stock options and stock awards granted to our directors and NEOs in 20202022 see the sections titled “Executive Compensation” and “Director Compensation.”
Limitation of Liability and Indemnification of Directors and Officers
Our amended and restated certificate of incorporation provides that no director would be personally liable to us or our stockholders for monetary damages for breach of fiduciary duty as a director, except as required by applicable
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law. For more information regarding the limitations of liability and indemnification see the section titled “Limitation of Liability and Indemnification.”
Related Party Transaction Policy
We have adopted a formal written policy that applies to our executive officers, directors, holders of more than five percent5% of any class of our voting securities and any member of the immediate family of, and any entity affiliated with, any of the foregoing persons. Such persons are not permitted to enter into a related-party transaction with us without the prior consent of our audit committee, or other independent members of our board of directors in the event it is inappropriate for our audit committee to review such transaction due to a conflict of interest. Any request for us to enter into a transaction with an executive officer, director, principal stockholder or any of their immediate family members or affiliates in which the amount involved exceeds $120,000 must first be presented to our audit committee for review, consideration and approval. In approving or rejecting any such proposal, our audit committee will consider the relevant facts and circumstances available and deemed relevant to our audit committee, including, but not limited to, whether the transaction will be on terms no less favorable than terms generally available to an unaffiliated third-partythird party under the same or similar circumstances and the extent of the related-party’srelated party’s interest in the transaction.
All of the transactions described in this section occurred prior to the adoption of this policy.
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CAUTIONARY STATEMENT REGARDING FORWARD-LOOKING STATEMENTS
This proxy statementProxy Statement contains forward-looking statements within the meaning of Section 27A of the Securities Act and Section 21E of the Exchange Act, which are subject to the “safe harbor” created by those sections and that represent the Company’s current expectations and beliefs. All statements other than statements of historical fact are “forward-looking statements” for purposes of federal and state securities laws and include, but are not limited to, statements of management’s expectations regarding the Company’s future compensation decisions, organization and performance. Forward-looking statements generally can be identified by the use of forward-looking terminology such as “may,” “will,” “should,” “expect,” “plan,” “anticipate,” “could,” “intend,” “target,” “project,” “contemplate,” “believe,” “estimate,” “predict,” “potential” or “continue” or the negatives of these terms or variations of them or similar terminology. The forward-looking statements and related assumptions involve risks and uncertainties that could cause actual results and outcomes to differ materially from any forward-looking statements or views expressed herein. The Company’s performance and the forward-looking statements contained herein are further qualified by a detailed discussion of associated risks set forth in the documents filed by the Company with the Securities and Exchange Commission. The forward-looking statements set forth the Company’s beliefs as of the date that such information was first provided, and the Company assumes no duty to update the forward-looking statements contained in this proxy statementProxy Statement to reflect any change except as required by law.
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OTHER MATTERS
Delinquent Section 16(A) Reports
Section 16(a) of the Exchange Act requires our directors and officers and holders of more than 10% of our common stock to file with the SEC initial reports of ownership of our common stock and other equity securities on a Form 3 and reports of changes in such ownership on a Form 4 or Form 5. Directors and officers and holders of 10% of our common stock are required by SEC regulations to furnish us with copies of all Section 16(a) forms they file. To our knowledge, based solely on a review of our records and representations made by our directors, officers and beneficial owner of more than ten percent10% of our common stock, regarding their filing obligations, all Section 16(a) filing requirements were satisfied with respect to the fiscal year ended December 31, 2020,2022, other thanthe following late filings: (i) a Form 4 filings on December 18, 2020 for Lip-Bu Tan, WIIG Communications Management LLC (and entities managed thereby), Walden Riverwood GP, LLC (and entities managed thereby), WRV GP II, LLC (and entities managed thereby),each of Kurt Wood, Stuart Merkadeau, Keith Breinlinger and Eric Hobbs for the withholding of shares to satisfy tax withholding obligations in connecting with the time vesting of restricted stock units, (ii) a Form 4 each of Siddhartha Kadia and James Paul McClaskey for a grant of restricted stock units and a grant of stock options, (iii) a Form 4 for each Eric Hobbs and Gregory Lucier for grant of restricted stock units, (iv) a Form 4 for James Paul McClaskey for a grant of restricted stock units, (v) a Form 3 for James Paul McClaskey and (vi) a Form 4 for each of Michael Moritz, John Chiminski, James Rothman, Jessica Hopfield, Igor Khandros, Ph.D.Gregory Lucier and Michael Marks, with respect to their salesElizabeth Nelson for a grant of sharesrestricted stock units and a grant of our common stock upon the closing of our underwritten secondary public offering on December 10, 2020.options.
Fiscal Year 20202022 Annual Report and SEC Filings
Our financial statements for our fiscal year ended December 31, 20202022 are included in our Annual Report on Form 10-K filed with the SEC on March 12, 2021February 23, 2023 (File No. 001-39388). This proxy statementProxy Statement and our Annual Report are posted on the Company’s website at https://investors.berkeleylights.com/investors.phenomex.com/ in the “Financial Information”“Financials” section of the Investors Relations webpage and are available from the SEC at its website at www.sec.gov. You may also obtain a copy of our Annual Report without charge by sending a written request to Berkeley Lights,PhenomeX Inc., Attn: Corporate Secretary, 5858 Horton Street, Suite 320, Emeryville, California 94608.
Company Website
We maintain a website at www.berkeleylights.com.www.phenomex.com. Information contained on, or that can be accessed through, our website is not intended to be incorporated by reference into this proxy statement,Proxy Statement, and references to our website address in this proxy statementProxy Statement are inactive textual references only.
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PROPOSALS OF STOCKHOLDERS FOR 20222024 ANNUAL MEETING
To be considered
Proposals Pursuant to Rule 14a-8. Pursuant to Rule 14a-8 under the Exchange Act, stockholders may present proper proposals for inclusion in the Proxy Statement and for consideration at our next year’s proxy materials,annual meeting of stockholders. To be eligible for inclusion in our Proxy Statement for the 2024 Annual Meeting, your proposal must be submitted in writingreceived by November 26, 2021, to our the attention of ourCorporate Secretary at Berkeley Lights, Inc.,our principal executive offices at 5858 Horton Street, Suite 320, Emeryville, CA 94608; provided94608, no later than December 16, 2023, and must otherwise comply with Rule 14a-8. While our board of directors will consider stockholder proposals, we reserve the right to omit from the proxy statement stockholder proposals that ifwe are not required to include under the date of the annual meeting is more than 30 days from May 14, 2022, theExchange Act, including Rule 14a-8.
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deadline is a reasonable time before we begin to print and send our proxy materials for next year’s annual meeting.Proposals Pursuant to theOur Bylaws. Under our bylaws, in order to bring any other business before the stockholders at the 2024 Annual Meeting, other than proposals pursuant to Rule 14a-8, you must notify us in writing and such notice must be received by us no earlier than January 25, 2024 and no later than February 24, 2024. You must comply with specific procedures set forth in our bylaws and the proposal must contain the specific information required by our bylaws. You may write to our Corporate Secretary at our principal executive offices at 5858 Horton Street, Suite 320, Emeryville, CA 94608, to deliver the notices discussed above and to request a copy of the relevant bylaw provisions regarding the requirements for making stockholder proposals pursuant to the bylaws.

If a stockholder who has notified us of his, her or its intention to present a proposal for next year’sat an annual meeting other than proposalsdoes not appear to be included in the proxy statement as described above,present his, her or to nominate a director, you must do so between January 14, 2022 and February 13, 2022; provided that if the date of thatits proposal at such annual meeting, is more than 30 days beforewe are not required to present the proposal for a vote at such annual meeting. We reserve the right to reject, rule out of order or more than 60 days after May 14, 2021, you must give noticetake other appropriate action with respect to any proposal that does not later thancomply with the 90th day prior to the annual meeting daterequirements of our amended and restated bylaws or if later, the 10th day following the day on which public disclosure of the annual meeting date is first made. You are also advised to review our bylaws, which contain additional requirements about advance notice of stockholder proposals and director nominations.other applicable requirements.
Nomination of Director Candidates
You may propose director candidates for consideration by our nominating and corporate governance committee by providing the information required by our amended and restated bylaws. Any such recommendations should include, among other things outlined in our amended and restated bylaws, the nominee’s name and qualifications for membership on our board of directors and should be directed to our Corporate Secretary at the address set forth above.

In addition, our amended and restated bylaws permit stockholders to nominate directors for election at an annual meeting of stockholders. To nominate a director, the stockholder must provide the information required by our amended and restated bylaws. In addition to satisfying the stockholderforegoing advance notice requirements under the Company’s bylaws, to comply with the universal proxy rules under the Exchange Act, stockholders who intend to solicit proxies in support of director nominees other than the Company’s nominees must give timelyprovide notice to our Secretary in accordance with our amended and restated bylaws, which, in general, require that sets forth the notice be receivedinformation required by our Secretary withinRule 14a-19 under the time period described above.Exchange Act no later than March 26, 2024.
Availability of Bylaws
A copy of our amended and restated bylaws may be obtained by accessing our public filings on the SEC’s website at www.sec.gov. You may also contact our Corporate Secretary at our principal executive office for a copy of the relevant bylaw provisions regarding the requirements for making stockholder proposals and nominating director candidates.
Our board of directors does not know of any matter to be brought before the 20212023 Annual Meeting other than the matters set forth in the Notice of 20212023 Annual Meeting of Stockholders and matters incident to the conduct of the 20212023 Annual Meeting. If any other matter should properly come before the 20212023 Annual Meeting, the persons named in the enclosed proxy card will have discretionary authority to vote all proxies with respect thereto in accordance with their best judgment. It is important that your shares of common stock be represented at the 20212023 Annual Meeting, regardless of the number of shares that you hold. You are, therefore, urged to vote by telephone, by using the Internet or by mail at your earliest convenience as instructed on the Notice of Internet Availability of Proxy Materials.
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By order of the Board of Directors,
erichobbssignatureforsecfia.jpg
Eric Hobbs,siddharthasignaturea.jpg
Siddhartha Kadia, Ph.D.
Chief Executive Officer and Director
March 31, 2021April 14, 2023
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